V PATHOGENIC DISEASE/PEST REACTION

For disease/pest reaction gene guidelines see section III. 8.

Note: In listings of multiple alleles, the chromosomes locations and ma: citations with generally be given with the particular allele that was located of mapped.

Reaction to Barley Yellow Dwarf Virus
Bdv1 {1363,1379}. 7D {1379}, 7DS {1363}. i: Jupeteco 73R (compared to Jupeteco 73S) {1363}.
v: Anza {1379}; Condor BW3991 {1379}; Tyrant BW3872 {1379}; Hahn BW4097 {1379}; Parrot BW10817 {1379}; Siren BW18643 {1379}; Many CIMMYT genotypes. Bdv1 is completely linked with Ltn, Lr34 and Yr18. See Ltn, Lr34, Yr18.

Note: BW = CIMMYT bread wheat accession number.
 
Bdv2 {058}. 7DL= T7DS.7DL-7Ai#1L {552}. tr: TC14 {059}.
  7D = T7DS-7Ai#1S.7Ai#1L {552}. tr: TC5, TC6, TC8, TC9, TC10.
  1B = T1BS-7A#1S.7Ai#1L {552}. tr: TC7 {447}.
7Ai#1s {552}. su: TAF2 {059}; Lines
5395 & 5395-243AA {552}.

Reaction to Cochliobolus sativus Ito & Kurib.
Disease: Cochliobolus root rot.
 
Crr {764}. Recessive. 5BL {764,765}. v: Apex {764}; Cadet {765}.

Reaction to Diuraphis noxia {Mordvilko}
Insect pest: Russian aphid.
 
Dn1 {286}. 7D {1288}. v: PI 137739 {286}.
Dn2 {286}. 7DL{863}. v: PI 262660 {286,863}.
ma: XksuA1-7D - 9.8 cM - Dn2 {863}.

According to Saidi & Quick {1250}, Dn1 and Dn2 are probably allelic. Reference stocks with each gene showed allelism with a gene in PI 262605.

Dn3 {1086}. Recessive. v: T. tauschii SQ24/T. turgidum TD65 {1086}. dv: T. tauschii SQ24 {1086}.

Dn4 {1250}. 1DL {863}. v: CORWA1 {260}; CI2401 {260}; PI151918 {260}; v: PI 372129 {1250}. ma: Xabc156-1D -11.6 cM - Dn4 {863}.

Dn5 {1249}. 7D {259}, 7DL {287}. i: Palmiet derivative 92RL28 {287}. v: STARS -
9302W-sib {259}. v: PI 294994 {259}.

Dn6 {1250}. v: CI 6501 {260}; PI 243781 {1250,1249}.

Reaction to Erysiphe graminis DC.
Disease: Powdery Mildew.

Note: Chancellor, used as a susceptible genetic background, for some near-isogenic lines probably carries Pm10 and Pm15 {1479}.

Pm1 {130}. [Mlt {1175}, Mla {348}]. 7A {1293}, 7AL {1305}. i: CI 14114 = As II/8*Chancellor {132}; Axminster/8*Chancellor {132}; CI 13836/8*Chancellor {132}; Kenya C6041/5*Federation {1168}; Norka/8*Chancellor {132}. s: CS*5/Axminster 7A {1293}. v: Anfield {098}; As II {130}; Axminster {130,1175}; Birdproof {165}; Bonus {1554}; Converse {1175}; CI 13836 {130}; Fedka {939}; Festival {1554}; Fram I {130}; Huron CI 3315 {1175,1554}; Kenora {1554}; Kenya W744 = C6041 {130,1175}; Norka {130,1175}; Pika {130}; Sweden W1230 {1554}; Thew {1175}; TU 4 {130}; Zhengzhou 871124 {570}. Sappo Pm2 Pm4b (Carries Lr20 {310}); Solo Pm2 Pm4b {052}. Mephisto Pm2 Pm9 {540}; Normandie Pm2 Pm9 {165}. BGRC 44514 Pm3a {1628}. Anfield Pm9 {1287};Pompe Pm9 {1287}; Ring Pm9 {1287}. ma: Co-seg. with Xcdo347-7A using NILs {864}; Co-segregation or close linkage with three RAPDs; one RAPD converted to a STS {570}.
Note: In Solo, Pm1 is translocated to chromosome 7D {052}.

Variation among Pm1 alleles was proposed as follows:
 
Pm1a {562}. [Pm1 {562}]. v: See earlier lists.
Pm1b {562}.   v: MocZlatka {562}.
Pm1c {562}. [Pm18 {853,562}]. v: M1N {1628,562}; In {540} M1N was described as an undesignated subline of Weihenstephan M1.
Pm1d {562}.   v: T. spelta var duhamelianum TRI2258 {562}.

Pm2 {130}. [Mlu {1175}, Mlx {1088}]. 5D {1007}, 5DS {945}. i: CI 14119 = CI 12632/8*Chancellor {132}; Federation*4 /Ulka {1168}; CI 14118 = Ulka/8*Chancellor {132}. v: Avalon {096}; Bounty {096}; Fenman {096}; Galahad {1531}; H8810/47 {130}; Synthetic ; Longbow {1531}; Maris Beacon {1592}; Maris Nimrod {1592}; Maris Sportsman {096}; Maris Templar {1592}; Norman {096}; Orestis {1079}; PI 92378 {1168}; PI 181374; {1168}; Sea Island {130}; Sentry {096}; S2303 {945}; TP 114/2*Starke deriv. {626}; Ulka {130,1175}; (Iumillo/T. tauschii) {1168}. XX186 = T. durum ‘Santa Marta’/Aeg. squarrosa BGRC 1458 Pm19 {853}; Compal Pm4b {854}; Timmo Pm4b {096}. Brigand Pm6 {096}; Brimstone Pm6 {1531}; CI 12632 Pm6 {130}; CI 12633 Pm6 {133}; Gawain Pm6 {1531}; Heiduck Pm6 {541}; Hustler Pm6 {096}; Kinsman Pm6 {096}; Mardler Pm6 {096}; Maris Fundin Pm6 {096}; Maris Huntsman Pm6 {1592}; TP 114 Pm6 {626}; Virtue Pm6 {096}. Hornet Pm8 {1531}. Solo Pm1 Pm4b {052}. Mephisto Pm1 Pm9 {540}; Normandie Pm1 Pm9 {165}. Rendezvous Pm4b Pm6 {1531}; Walter Pm4b Pm6 {1428}. Apollo Pm4b Pm8 {541}. Crossbow Pm5 Pm6 {098}; Parade Pm5 Pm6 {1531}. Halle Stamm 13471 Mld {097}; Maris Dove Mld {1592}. dv:Aeg. squarrosa BGRC 1458 {853}. Forty accessions of T. tauschii {852}. ma:Pm2 - 3.5cM - Xbcd1871-5D using F2s {864}.

Pm3

Pm3a {130,132}. [Mla {1168}]. 1A {1007}, 1AS {943,947}.i: Asosan/8*Chancellor {132} = CI14120. Asosan / 3*Federation {1168}. v: Asosan {130,1168}; BGRC 44514 Pm1 {1628}; Coker 797 {786}; Florida 301 {786}; Florida 302 {786}; Hadden {097}; Halle Stamm {097}; PI 46890 {1439}; Norin 3 {1134}; Norin 29 {1134}; Saluda {786}; Tyler {1419}. ma:Xbcd1434-1A - 1.3 cM - Pm3 using NILs {864}; Xwhs179-1A - 3.3 cM - Pm3 {522}.

Pm3b {130,132}. [Mlc {165}]. 1A {1007}. i: Chul*8/Chancellor {132}; T. sphaerococcum*8/Chancellor = CI 15887 {539}. v: Chul {165}. ma: Xbcd1434-1A - 1.3 cM - Pm3b using NILs {864}.

Pm3c {130,132}. [Mls {1175}]. 1A {134,1007}. i: Sonora/8*Chancellor {132} = CI 14122; Sonora/4* Federation {1168}; Triticale/8*Chancellor {539}. s: CS*7/Indian 1A {134}. v: Borenos {854}; Cawnpore {1628}; CI 3008 {130}; CI 4546 {130}; Hindukusch {1628}; Indian {1175}; Sonora {130,1168}; Sturgeon {1175}.

Pm3d {1628}. [Ml-k {540}], Mlk {434}. 1A {1628}. v: Kolibri {540,542,1628}; Herold {540}; Ralle {540}; Socrates {heterogeneous} {540}; Star {heterogeneous} {540}; Syros {540}. Kadett Pm4b {540}; Turbo Pm4b {540}.

Pm3e {1628}. v: Sydney University Accession W150 =AUS 6449 {939,1628}.

Pm3f {1628}. i: Michigan Amber/8*Chancellor {1628}. This allele was distinguished from Pm3c with only one of 13 pathogen cultures.

Pm4 [{131}].

Pm4a {1464}. [Pm4 {131}]. 2AL {1464}. i: CI 14123 = Khapli/8*Chancellor {131}; CI 14124 = Yuma/8*Chancellor {131}. v: Steinwedel*2/Khapli {939}. tv: Khapli {131}; Valgerado {097}; Yuma {131}. ma: Co-seg with Xbcd1231-2A.2 & Xcdo678-2A using F2s {864}; Xbcd1231-2A.1 - 1.5cM - Pm4 - 1.56cM - Xbcd292-2A {864}.

Pm4b {1464}. [Mle {1591}]. 2A {052}, 2AL {1464}. i: Federation*7/T. carthlicum W804 {1464}. v: Achill {540}; Ajax {540}; Arkas {540}; Armada {096}; Botri (heterogeneous) {854}; ELS {1591}; Facta {854}; Factor (heterogeneous) {854}; Fakon {854}; Fazit {854}; Hermes {540}; Horizont {540}; Maris Halberd; Max {540}; Olymp {540}; Orbis {540}; S-25 {052}; Ronos {1079}; S-28 {052}; TP 229{626,1591}; Weihenstephan M1 {1591}; VPM1 {097}. Rang Pm1 {052}. Solo Pm1 Pm2 {052,540}. Compal Pm2 {854}. Rendezvous Pm2 Pm6 {1531}; Timmo Pm2 Pm6 {096}; Walter Pm2 Pm6 {1428}. Apollo Pm2 Pm8 {541,802}; Kadett Pm3d {540}; Turbo Pm3d {540}. Boxer Pm5 {541}; Mission Pm5 {78,541,1531}. Sorbas Pm6 {541}. Kronjuwel Pm8 {541}.

Pm5 {787}. [Mli {540,558}], mlH {771}. Recessive. 7B{964}, 7BL {771}. i: Hope/8* Chancellor = CI 14125 {570}. s: CS*6/Hope 7B {771,964}. v: Alidos {854}; Aotea {964}; Caldwell {786}; Ga 1123 {786}; Glenwari {964}; Hardired {786}; Hope {964}; H-44 {964}; Kontrast {854}; Kormoran {1079}; Lawrence {964}; Redcoat {097}; Redman {964}; Renown {964}; Selpek {540}; Sicco {096}; Spica {964}; Warigo {964}. Arthur Pm6 {786}; Coker 983 Pm6 {786}; Double Crop Pm6 {786}. Granada Pm8 {541}; Sensor Pm8 {541}. Wheats identified originally as having Mli are listed below. Pm5 was transferred to hexaploid wheat fromT. dicoccum via Hope and H-44, but there is not apparent relationship of the Mli group with this source.

v: Aquila {096,541}; Carimulti {541}; Cariplus {541}; Dolomit {541}; Falke {541}; Flanders {096}; Ibis {096}; Kormoran {541}; Kraka {541}; Markant {541}; Mercia {1531}; Milan {541}; Reiher {541}; Rektor {541}; Rothwell Perdix {096}; Severin {541}; Sicco {096}. Sperber {541}; Tukan {541}; Urban {541}; Wattines {541}. Boxer Pm4b {541}; Mission Pm4b {541,1531}. Bert Pm6 {541}. Kristall Pm8 {541}. Crossbow Pm2 Pm6 {098}; Parade Pm2 Pm6 {1531}.

Pm6 {627}. [Mlf {626}]. 2B {1088}. v: Abe {097,1256}; Coker747 {786,1079}; Timgalen {098}; 1969 IVGS Line C {626}; Mengavi {097}; TP 114/2*Starke deriv. B {626}. Oasis {786}. Brigand Pm2 {096}; Brimstone Pm2 {1531}; CI 12632 Pm2 {626,1088}; CI 12633 Pm2 {626,1088}]; Garwain Pm2 {1531}; Heiduck Pm2 {541}; Hustler Pm2 {096}; Kinsman Pm2 {096}; Mardler Pm2 {096}; Maris Fundin Pm2 {096}; Maris Huntsman Pm2 {1592}; TP 114 Pm2 {626}; Virtue Pm2 {096}. Rendezvous Pm2 Pm4b {1531}; Timmo Pm2 Pm4b {096}; Walter Pm2 Pm4b {1428}. Crossbow Pm2 Pm5 {098}; Parade Pm2 Pm5 {1531}. Sorbas Pm4b {541}. Arthur Pm5 {097,786}; Double Crop Pm5 {786}.

Pm7. Derived from S. cereale cv. Rosen. 4BL {270,271,389} = T4BS.4BL-5R {543}; more recently revised to T4BS.4BL-2R#IL {380,389}. i: = Transec/8*Chancellor v: Transfed {269}; Transec {273}.

T4BS.4BL-5RL {543}, but more recently shown to be T4BS.4BL-2RL {380}.

Pm8. 1R{1B}, 1BL.1RS. v: Derivatives of Petkus rye - see Yr9, Lr26, Sr31. Ambassador {1531}; Corinthian {1531}; Dauntless {1531}; Disponent {541}; GR876 {753}; Halle Stamm {097}; Hammer {098}; Slejpner {1531}; ST1-25 {201}; Stetson {1531}; Stuart {096}. Hornet Pm2 {1531}. Apollo Pm2 Pm4b {541}. Kronjuwel Pm4b {541}. Granada Pm5 {541}; Kristall Pm5 {541}. Sensor Pm5 {541}. Others {1208}.

Crosses between three lines with Pm8 and Helami-105a 1BL.1RS line with Pm17, indicated that Pm8 and Pm17 were allelic {524}. Earlier, these genes were reported to be genetically independent {1463}. tv: Cando*2/Veery = KS91WGRC14 {381}.

Suppressor of Pm8:
Some wheats which, on the basis of cytological and rust tests carry 1RS from Petkus rye, do not express resistance due to presence of a suppressor {385}. Zeller & Hsam {1625} located a suppressor of Pm8 and Pm17 in chromosome 7D of Caribo. Mildew resistance was suppressed in Florida, Heinrich, Ikarus, Olymp and Sabina, which are derivatives of Caribo with 1BL.1RS. According to Ren et al. {1209}, SuPm8 does not suppress Pm17. Hanusova et al. {492} listed 16 wheats that carry a suppressor of Pm8; 111 wheats did not carry the suppressor. In contrast, a high frequency of suppression occurred in CIMMYT wheats {108,1208}. Further genotypes are identified in {491}.
SuPm8 {1209}. 1AS {1209}. v: Wheats with Gli-A1a {1209}. Lists in {108,491,1208}.

Pm9 {347}. 7A {347}, 7AL. v: Anfield Pm1 {1287}, Pompe Pm1 {1287}, Ring Pm1 {1287}. Mephisto Pm1 Pm2 {540}; Normandie Pm1 Pm2 {347}.

Pm10 {1482}. 1D {1482}. v: Norin 4 {1482}; Norin 26 {1482}; Norin 29 {1482}; Penjamo 62 {1482}; Shin-chunaga {1482}. T. spelta duhamelianum Pm11 {1481}. Pm10 was detected using a culture derived from a hybrid of E. g. tritici and E. g. agropyri.

Pm11 {1481}. 6BS {1481}. v: Chinese Spring {1481}; Salmon {1481}; T. compactum No. 44 {1481}. T. spelta duhamelianum Pm10 {1481}.
Pm11 was detected using a culture derived from a hybrid of E. g. tritici and E. g. agropyri.

Pm12 {1017}. Derived from Ae. speltoides. 6B = 6BS-6SS.6Sl {598,572}. The earlier location of 6A {1017} was not correct. v: Wembley*6/Ae. speltoides, #31 {1017,598}. 6SlS {598}. al: Ae. speltoides CL214008 = K {1017}. ma:Pm12 was mapped to a translocated 6SlS segment proximal to Xpsr551-6B {598}.

Pm13. Derived from T longissimum. 3B {173}, = T3BL.3BS-3S1#1S {389}. v: T. longissimum derivative R1A {174}. 3D {173} = T3DL.3DS-3Sl#1S {389}. tv: T. ongissimum derivative R1D {174}. 3Sl#1S. al: T. Longissimum.

Pm14. 6B {1478}. v: Akabozu Pm15 {1478}; Kokeshi-komugi Pm15 {1478}; Norin 10 Pm15 {1478}.
Pm14 and Pm15 were detected using hybrids between E. g. tritici and E. g. agropyri cultures.

Pm15. 7DS {1478}. v: Norin 4 Pm10 {1478}; Norin 26 Pm10 {1478}; Shin-chunaga Pm10 {1478}; T. machasubletschumicumPm10 {1478}. Chinese Spring Pm11 {1478}; T. compactum No. 44 Pm11 {1478}. Akabozu Pm14 {1478}; Kokeshi-komugi Pm14 {1478}; Norin 10 Pm14 {1478}.

Pm14 and Pm15 were detected using hybrids between E. g. tritici and E. g. agropyri cultures.

Pm16 {1201}. 4A {1201}. v: Norman lines with resistance from T. dicoccoides CL1060025 {1201}. tv: T. dicoccoides CL1060025 {1201}.

Pm17 [{097,838}] {544}. 1AS = T1AL.1R#2S {1624,185,389}. v: Amigo. 1BS = T1BL.1R#2S {561}. v: Century {216}. v: Helami 105 Pm5 {561}; TAM107 {216}; TAM200 {216}; TAM201 {216}.

Pm8 and Pm17 were reported to be allelic {524}.

Suppressor of Pm17
A suppressor of Pm17, as well as of Pm8, was located in chromosome 7D of Caribo {1625}. Gene SuPm8 identified by Ren et al. {1209} did not suppress Pm17.

Pm18. Deleted, see Pm1c.

Pm19 {853}. 7D {853}. v: Synthetic XX186 Pm2 {853}. T. durum ‘Moroccos 183'/Aeg. squarrossa AE 457/78 {853}. dv: Ae. squarrosa {853}.

Pm20 {386}. [M1P6L {543}]. 6BL = T6BS.6R#2L {543,386,389}. v: KS93WGRC28 = PI 583795 {386,382}. 6RL. su: 6R{6D} {543}. ad: 6R addition {543}. al: Prolific rye {543}.

Pm21 {1177}. 6AS = T6AL.6VS {1177}. v: 9 independent translocations {1177}. ma: RAPD OPH171900 (synonym 'OPH17-1900') was associated with Pm21 and RAPD OPH171000 (synonym OPH17-1000') with its absence {1176}.

Pm22 {1134}. 1D {1134}. v: Elia {1134}; Est Mottin {1134}; Ovest {1134}; Tudest {1134}; Virest {1134}.

It was not possible to test Pm22 for possible allelism with Pm10.

Pm23 {1618}. 5A {1618}. v: Line 81-7241 Pm8 {1618}.
Although Line 81-7241 carries Pm8, evidence was presented to indicate that it was suppressed in Line 81-7241 and, by inference, also in Chinese Spring.

Pm24 {571}. 6D {571}. v: Chiyacao {571}.

Complex genotypes: Drabent {heterogeneous} Pm2 Pm4bPm9/Pm1 Pm2 Pm4b Pm9 {1287}; Nemares Pm1 Pm2Pm4b Pm6 Pm9 {1287}; Planet, Sappo & Walter Pm1 Pm2 Pm4b Pm9 {096,097,540,1287,1428}.

Lists in {1629} (French wheats), {1608,572} (Chinese wheats), {1101} (Western Siberian wheats).

Temporary designations:

Ml-Ad {854}. v: Adlungs Alemannen {854}.

Mlar {854}. v: Abo {1629}; Aristide {1629}; Courtot {1629}.

Ml-Br {854}. v: Bretonischer Bartweizen {854}.

Mld {096}. 4B {097}. v: Halle 13471 Pm2 {096}. H8810/47 Pm2 {096}; Maris Dove Pm2 {096}. tv: T. durum line {096}.

Ml-Ga {854}. v: Garnet {854}; many old German cultivars {854}.

mlre {1220}. v: RE714 {1220}.

PmTmb {1343,1344}. v: NC94-3778 {1344}. dv:T. monococcum PI 427662 {1344}. ma: Associated with 3 RAPDs {1344}.

Reaction to Fusarium graminearum
Disease : Fusarium head scab (= Fhs).

Fhs1 {1096}. v: Line A {1096}. Ning 7840 Fhs2 {1097}.

Fhs2 {1096}. v: Line B {1096}. Ning 7840 Fhs1 {1096}.

Reaction to Heterodera avenae Woll.
Cereal root eelworm; cereal cyst nematode.

Cre1. [Cre {1388}]. 2B {1388}. i: AP = Prins*8/AUS10894 {1579}. v: Loros CI 3779 {1388}; AUS 10894 {1056}. ma: Xglk605-2B - 7.3cM - Cre1 - 8.4cM - Xcdo588-2B/Xabc451-2B {1579}.

Cre2 {238}. Derived from Ae. ventricosa {238}. v: H93-8 {238}.

Cre3. [CcnD1 {329}, Ccn-D1 {328}]. 2DL {328}. v: Synthetic hexaploids {329}. dv: T. tauschii accessions AUS 18912 {328}; AUS 18913 {328}; CPI 110809 {329}; CPI 110810 {328}.

Cre4. [CcnD2 {329}, Ccn-D2 {328}]. 2D {328}. dv:T. tauschii accessions AUS 18914 {329}; CPI 110813 {328}.

Reaction to Mayetiola destructor (Say) (Phytophaga destructor) (Say)
Insect pest: Hessian fly.

H1 {1087}. i: Dawson/3*Poso, 6179 {1087}. v: Big Club 43 H2 {1441}; Dawson H2 {166,1087}; Poso 42 H2 {1441}.

H2 {1087}. i: Dawson/3*Poso, 6232 {1087}. v: Big Club 43 H1 {1441}; Dawson H1 {166,1087}; Poso 42 H1 {1441}.

H3 {156}. Recessive. 5A {425,1105}. i: Carol = Newton-207*5/Larned {1107}. v: Ace {426}; Arthur {426}; Becker {749}; Cardinal {750}; Dual {1273}; Frankenmuth {341}; Georgia 1123 {426}; GR855 {751}; GR876 {753}; Ionia {426}; Larned {824}; Logan {426}; Monon {157}; Norkan {904}; Ottawa {547}; Purdue B 36162 A13-12 {156}; PI 468960 {1439}; Redcoat {1273}; Reed {1273}; Riley {1273}; Roland {148}; Russell {426}; Shawnee {547}; Titan {747}; Todd {426}; W38 {156}. Clara Fay H6 {375}. ma: Cosegregation of H3 and a RAPD {296}.
Allen et al. {019} considered that H3 and H4 may be allelic. Also suggested by Shands and Cartwright {1317}. Linkage of 10.5 + 2% involving H3 and Pm3a in PI 468960 was attributed to a chromosome 1A/5A translocation {1437}.
H4. [h4 {1441}]. Recessive. v: Dixon {1441}; Java {1441}.

H4 confered resistance to race A, but not to race B.

H5 {1317}. 1AS {1222}. Temperature sensitive {1413}. v: Abe {162}; Arthur 71 {162}; Beau {875}; Downy {1223}; Oasis {1109}; Ribeiro {1317}; Sullivan {1110}. tv: Giorgio 331-4 {1090}; PI 94567-6 {1317}; PI 94571-14 {1317}. ma: Cosegregation of H5 and two RAPDs {296}.

H6 {019}. 5A {425}. i: Erin = Newton-207*7/Arthur 71 {1107}; Flynn = Newton-207*7/Knox 62 {1107}. v: Adder {1319}; Benhur {426}; Caldwell {1421}; Compton {1318}; CI 12855 {019}; Excel {752}; Fillmore {1106}; Knox 62 {426}; Lathrop {426}. Clara Fay H3 {375}. tv: Purdue 4835 A4-6 {1105}. PI 94587 H11 H16 {019}. ma: Cosegregation with three RAPDs {296}.

H7 & H8 {425}. Duplicate factors. H7 is located in chromosome 5D {026}. v: Adena {748}; Seneca {026,425}.

H9 {1420}. 5A {162}. i: Iris = Newton-207*7/Ella {1107} v: Ella {875}; Line 822-34 {162}. Elva CI 17714 H10 {162}; Line 812-24 H10 {1421}; Line 817-2 H10 {1421}; Stella H10 {875}. ma: Cosegregation with two RAPDs {296}.

H10 {1104}. 5A {162}. i: Joy = Newton-207*3/IN76529A5-3-3 {1107} v: IN 76529 {875}. Elva CI 17714 H9 {162}; Line 817-2H9 {162}; Stella H9 {875}. ma: Cosegregation with one RAPD and close linkage to another RAPD {296}.

H11 {1422}. 1A {1222}. i: Karen = Newton-207*4/IN916-1-3-1-47-1 {1107} v: Kay {875,375}; Line 916 {1422}; Line 920 {1422}; Line 941 {1422}. tv: T. turgidum PI 94587 H6 H16 {1422}. ma: Close linkage with two RAPDs {296}.

H12 {1092}. 5A {1098}. i: Lola = Newton-207*4/Luso {1107}. v: Luso {1092}. ma: Cosegregation with one RAPD and close linkage of H12 to another RAPD {296}.

H13 {1104}. 6DL {441}. i: Molly = Newton-207*7/3/KU221-19/Eagle/ KS806 {1107} v: KS81H1640HF {441}; T. turgidum var. durum cv. Gulab KU 134/T. tauschii KU 2076, KU 221-14 {525}; T. turgidum var. persicum straminium KU 138/T. tauschii KU 2076, KU221-19 {525}. dv: T. tauschii KU 2076 {525}. ma: Cosegregation with a RAPD {296}.

H14 {875}. 5A {875}. tv: IN 81601A2-3-3 {875}. ELS 6404-160 H15 {875}. ma: Cosegregation with a RAPD {296}.

H15 {875}. 5A {875}. tv: IN 81602C5-3-3 {875}. ELS 6404-160 H14 {875}.

H16 {1106}. 5A {1098}. tv: IN 80164H5-2-9 {1106}. PI 94587 H6 H11 {1106}. tv: IN80164 {1097}. ma: Cosegregation of H16 and a RAPD {296}.

H17 {1090}. 5A {1090}. tv: PI 428435 {1090}. ma: Cosegregation of H17 and a RAPD {296}.

H18 {1090}. v: Marquillo {426,874}; Shield {198}.

H19 {1089}. tv: PI 422297 {1089}. This wheat possesses a second gene which is allelic or closely linked with H16 {1089}. tv: IN84702 {1097}. PI422297 H29 {1097}.

H20 {025}. 2B {025}. tv: Jori {025}.

H21. 2B {383} = 2BS.2R#2L {389}. v: Hamlet = KS89WGRC8 {1312}; KSWR 69-2-4-3 {383}; KS85HF 011-5 {383}. ad: KSWR 297h-1-1-9 {383}. al: Chaupon rye {383}.

H22 {1199}. 1D {1199}. v: KS86WGRC1 {1199}.

H23 {1199}. 6D {442}, 6DL {1199}. v: KS89WGRC3 {442}. ma: H23 - 6.9 cM - XksuH4-6D {861}.

H24 {1199}. 3D {442,1199}, 6DL {861}. v: KS89WGRC6 {442}. ma: H24 - 5.9 cM - Xbcd451-6D/Xcdo482-6D {861}.

H25 6B {384} = T6BS.6BL-6R#1L {389}. v: 88HF16 = WGRC17 {384}.

4B {384} = T4BS.4BL-6R#1L {389}. v: 88HF79, 88HF80 = WGRC18, 88HF81, 88HF117 = WGRC19 {384}. 4A {384} = Ti4AS.4AL-6R#1L-4AL {389}. v: 89HF17, 89HF18, 89HF25, 88HF32, 88HF51, 88HF89 = WGRC20 {384}. 6R {384}. al: Balbo rye {384}.

H26 4D {217}. v: KS92WGRC26 {217}. dv: T. tauschii TA2473 {217}.

H27 {235}. 4Mv {235}. su: H-93-33 {235}. al:Ae. ventricosa No. 10 {235}; Ae. ventricosa No. 11 {235}.

H28 {171}. 5A {171}. tv: PI59190 {171}.

H29 {1095}. [H27 {171}]. 5A {1097}. tv: PI422297 H19 {1097}.

A recombination value of 12.0% between leaf-rust reaction {possibly Lr10} and Hessian-fly reaction in Selection 52400 was reported {018}.

Reaction to Meloidogyne spp.
Disease: Root rot nematode, root knot eelworm

Rkn {632}. dv: A. squarrosa G3489. v: Prosquare, a synthetic hexaploid of Produra/A. squarrosa G3489 {632}.

Rkn-mn1 {1621}. 3B {590}. v: Chinese Spring/Ae. variabilis No1//Rescler/3/Lutin {1620}.

Reaction to Mycosphaerella graminicola {Fuckel} Schroeter
Disease: Septoria tritici blotch

Stb1. [Slb1 {1586}]. v: Bulgaria 88 {1586}; Oasis {1586}; Sullivan {1586}.

Stb2. [Slb2 {1586}]. v: Nova Prata {1586}; Veranopolis {1586}.

Stb3. [Slb3 {1586}]. v: Israel 493 {1586}.

Stb4 {1410}. v: Cleo {1410}; Tadinia {1410}; Tadorna {1410}.

Stb4 segregated independently of Stb1 but its relationship with Stb2 and Stb3 is unknown.

Reaction to Phaeosphaeria nodorum
Disease: Septoria nodorum blotch

Snb1 {856}. 3AL {856}. v: Red Chief. EE8 Snb2.

Snb2 {856}. 2AL {856}. v: EE8 Snb1.

Snb3 {1594}. 5DL {1594}. s: CS*/Synthetic 5D {1594}. v: Synthetic {1594}. al: T. tauschii {1594}.

Temporary designation:

SnbTM {856,857}. 3A {857}, 3AL {856}. tv: T. timopheevii /2*Wakooma {856}; T. timopheevii PI 290518. T. timopheevii derivatives: S3-6 {857}, S9-10 {857}, S12-1 {857}.

Reaction to Pseudocercosporella herpotrichoides {Fron} Deighton
Disease: eyespot, strawbreaker footrot.

Pch {261}. 7D {591,592}, 7DL {708,1603}. s: Courtot*/Roazon 7D {592}; Hobbit Sib*/VPM1 7D {591}. v: Ae ventricosa derivative {261}; H-93-70 {236,1521}; Hyak {021}; Madsen {020}; Rendezvous {1603}; Roazon {591}; 5L 219 {1521}. al: Ae. ventricosa {261}.

Pch1 is closely linked with Ep-V1 {973}. Delibes et al. {236} concluded that Pch was not located in chromosome 7D whereas Law et al. {776} found that H-93-70 possessed a unique allele, Ep-D1b, in common with VPM1 and its derivatives. Eyespot resistance and Ep-A1b in chromosome 7A were genetically associated {704}.

Pch2 {228}. 7A {704}, 7AL {228,229}. s: CS*/Cappelle Desprez 7A {704,228}. v: Cappelle Desprez {704,228}. ma:Xcdo347-7A (distal) - 11 cM - Pch2 - 18.8 cM - Xwg380-7A (proximal){229}.

Pch3 {616}. ad: CS + 4V {1050}.
Temporary Designation:

PchDv {618}. 4V {618}. ad: Wheat + 4V {618}. s: Wheat 4V (group IV) {618}.