Note: Some near isogenic lines are based on Marquis. The genes present in the Marquis background are not listed for those NILs.

Sr1. Deleted - see Sr9d.

Sr2 {38,677}. Recessive allele. Adult plant response. 3BS {499}. s: CS*6/Hope 3B {499}. v: Warigo Sr7b Sr17 {499}. Suneca Sr8a Sr17 {485}. Hopps Sr9d {499}. Lancer Sr9d Sr17 {679}; Scout Sr9d Sr17 {679}. See also {499,1040}.

Sr2 is associated with pseudo-black chaff, {742,1102} and seedling chlorosis (see {149}) and occurs very frequently in commercial wheats, especially in germplasm produced and distributed by CIMMYT.

Sr3 & Sr4 {047}. v: Marquillo - based on early data. No stocks available.

Sr5 {047}. 6D {939,1308,1626}, 6DS {939}. i: I Sr5-Ra {828}; I Sr5-Rb {828}; Sr5/7*LMPG {685}; Thatcher/10*Marquis {686}. s: CS*6/Thatcher 6D {1308}. v: Admonter Früh {072}; Dacia {979}; Dong-Fang-Hong 2 {564}; Dong-Fang-Hong 6 {564}; Feng-Kong {563}; Hochzucht {046}; Hybrid 80-3 {072}; Jubilejna {068}; Juna {076}; Kanred {1308}; Ke-Fang 1 {564}; Stabil {072}; Viginta {071}; Vrakunski {072}. Victor Sr6 Sr8a {979}. Glenlea Sr6 Sr9b {327}. Qing-Chung 5 Sr6 Sr11 {564}. An-Hewi II Sr8a {564}; Erythrospermum 974 Sr8a {072}. N.P.789 Sr11 {1555}. Jing-Hong Sr17 {564}; Jing-Hong 2 Sr17 {564}. Amika Sr31 {076}; Dong-Xie 3 Sr31 {563}; Dong-Xie 4 Sr31 {563}; Istra Sr31 {076}; Solaris Sr31 {076}. Beijing 10 SrTmp {564}.

Sr6 {687}. [SrKa1 {1167}]. 2D {1293,1308,1577},2DS {942}. i: I Sr6-Ra {828}; Kenya 58/10*Marquis {468,675}. Sr6/9*LMPG {685}. s: CS*5/Red Egyptian 2D {1308}. v: Africa 43 {669}; Eureka {468,844}; Kenya stocks {669,670,673,687,689,1167,1557}; McMurachy {679}; Shield {198}. Fortuna Sr7a {679}; Kentana 52 Sr7a {678,1577}. Kiric 66 Sr7b {979}. Bowie Sr8a {1553}. Eurga Sr11 {1553}; Siete Cerros Sr11 {033}. Victor I Sr5 Sr8a {979}. Glenlea {heterogeneous} Sr5 Sr9b {327}. Lerma Rojo 64 Sr7b Sr9a {979}. Red Egyptian Sr8a Sr9a {687,1308}. No. 466 Sr9b Sr10 {689}. Gamut Sr9b Sr11 {1555}. See also {1553}.

Sr7 {830}. 4A {671,830,1293}, 4AL {939,1308}.

Sr7a {830}. [Sr7 {687}]. i: Egypt Na101/6*Marquis {468}; Kenya 117A/6*Marquis {468}; Sr7a/9*LMPG {685}. s: CS*7/Kenya Farmer 4B {830}; CS*8/Sapporo 4B {830}. v: Egypt Na101 {669}; Kenya stocks {669,670,673,687,689}; Sapporo Haru Komugi Ichigo {689}. Fortuna Sr6 {679}; Kentana 52 Sr6 {689}. French Peace Sr9a Sr13 {680}. Egypt Na95 Sr9b Sr10 {687}. W3746 Sr12 {1371}. Khapstein Sr13 Sr14 {674}.

Sr7b {830}. i: I Sr7b-Ra {828}. v: Warigo Sr2 Sr17 {499}. Kiric 66 Sr6 {979}. Roussalka Sr8a {979}. Red Bobs Sr10 {308}. Nell Sr17 {1565}; Spica Sr17 {939}. Marquis Sr18 Sr19 Sr20 {675,830}.

Sr8 6A {1293,1308}, 6AS {929,1368}.

Sr8a {1368}. [Sr8 {687}]. i: I Sr8a-Ra {828}; Red Egyptian/10*Marquis {686}; Sr8a/9*LMPG {685}. s: CS*5/Red Egyptian 6A {1308}. v: Marimp 3 {979}; Mentana {844}; Strampelli {979}. Hartog Sr2 Sr12 {127}. Suneca Sr2 Sr17 {485}. An-Hewi II Sr5 {564}; Erythrospermum 974 Sr5 {072}. Victor 1 Sr5 Sr6 {979}. Red Egyptian Sr6 Sr9a {687}. Roussalka Sr7b {979}. Frontana Sr9b {689}; Magnif G Sr9b{689}; Pitic 62 Sr9b {033}; Rio Negro Sr9b {689}. E-Gan-Zao Sr17 {564}; Golden Valley Sr17 {979}.

Sr8b {1368}. [SrBB]. v: Barleta Benvenuto {1368}; Klein Titan {1368}. Bezostaya Sr5{979}; Klein Cometa Sr30 {1368}.

Sr9 {676}. 2B {671,677,828,1308}, 2BL {944,946,951,1307,1582}.

Sr9a {676}. [Sr9 {687}]. i: I Sr9a-Ra {828}; Red Egyptian/10*Marquis {686}; Sr9a/9*LMPG {685}. s: CS*4/Red Egyptian 2B {1308}. v: Red Egyptian Sr6 Sr8a {687}. French Peace Sr7a Sr13 {680}. Excel Sr8a Sr17 {752}.

Sr9b {468}. [Sr9 {687}, SrKb1 {468}]. i: Kenya 117A/10*Marquis {686}; Sr9b/10*LMPG {685}. s: CS*7/Kenya Farmer 2B {939}. v: Gamenya {844}; Kenya stocks {669,670,673,687,689,1557}. Glenlea Sr5 Sr6 {heterogeneous} {327}. No. 466 Sr6 Sr10 {689}. Gamut Sr6 Sr11 {1555}. Egypt Na95 Sr7a Sr10 {636}. Frontana Sr8a {689}. Magnif G Sr8a {689}; Pitic 62 Sr8a {033}; Rio Negro Sr8a {689}. Veadeira Sr10 {687}. Robin Sr11 {879}. Festival Sr15 {1553}; Kenora Sr15 {1553}. See also {1553}.

Sr9c Originally reserved for SrTt1 - now Sr36.

Sr9d {678,831}. [Sr1 {047,676,677}]. i: Hope/10*Marquis {677}; H-44/10*Marquis {677}; I Hope 2B-Ra {828}. Sr9d/8*LMPG {685}. v: Hopps Sr2 {1040}. Lancer Sr2 Sr17 {679}; Scout Sr2 Sr17 {679}. tv: Arnautka {939}; Mindum {939}; Spelmar {939}.

Sr9e {951}. [Srd1v {642}], Srv{1391}. v:Vernstein {845}; SST 16 {1324} SST 33 {785}; SST 66 {785}; SST 3R {1324}; Sunstar Sr8a Sr12 {939}. Combination III Sr36 {841}. tv: Vernal emmer {1391}; CI 7778 {845}. Sr9e occurs in many tetraploid wheats {939,1378}.

Sr9f {826}. v: Chinese Spring {826}. Not present in the near-isogenic I Sr9a-Ra {826}.

Sr9g {965}. s: CS*7/Marquis 2B Sr16 {965}; CS*4/Thatcher 2B Sr16 {965}. v: Hochzücht Sr5 Sr12 {965}. Lee Sr11 Sr16 {965}. Celebration Sr12 Sr16 {965}. Eagle Sr26 {842}. See also {504}. tv: Acme {965}; Iumillo {965}; Kubanka {965}.

Sr10 {687}. 2B {686,939}. i: Egypt Na95/4*Marquis {468} v: Federation {939}; Geneva {1412}; Hazen {049}; Kenya stocks {669,670,673,687}. No. 466 Sr6 Sr9b {689}. Red Bobs Sr7b {308}. Egypt Na95Sr7a Sr9b {687}.

Sr11 {468}. [Sr11 and Sr12 {687}]. 6B {671,1143,1293,1309}, 6BL {1297}. i: I Sr11-Ra {828}; Lee/10*Marquis {686}. s: CS*7/Kenya Farmer 6B {830}; CS*9/Timstein 6B {1308}. v: Charter {844}; Flevina {072}; Gabo {687}; Kenya stocks {670,673,844,1557}; Sonora 64 {033}; Sylvia {071}; Timstein {687,1308}; Tobari 66 {033}; Yalta {844}. N.P.790 Sr5 {1555}. Qing-Chung 5 Sr5 Sr6 {564}. Eurga Sr6 {1553}. Gamut Sr6 Sr9b {1555}. Robin Sr9b {879}. Lee Sr9g Sr16 {687}. Prospect SrWld {197}. See also {1553}.

A resistance gene allelic with Sr11 was found in Chinese Spring {938}, but the P. graminis culture for its detection was lost.

Sr12 {1332}. Recessive. 3B {1332,682}, 3BS {968}. s: CS*7/Marquis Selection 3B Sr16 {1332}; CS*5/Thatcher 3B Sr16 {1332}. v: Marquillo {682}; Tincurrin {939}; Windebri {939}. W3746 Sr7a {1371}. tv: Postulated for several durums {1378}.

Sr12 is more widespread and probably more effective in conferring resistance than is usually acknowledged {939}.

Sr13 {674}. 6AL {929}. i: Khapstein/10*Marquis {686}; Sr13/9*LMPG {685}. v: French Peace Sr7a Sr9a {680}. tv: Khapli Sr14 {674}.

Sr14 {674}. 1BL {933}. i: Khapstein/10*Marquis {686}. v: Line A {933}. tv: Khapli Sr13 {674}.

Sr15 {1554}. 7A {1293,1554}, 7AL {1305}. v: Present in stocks possessing the Pm1 and Lr20 {931,1554}. See Reaction to Erysiphe graminis and Reaction to P. recondita.

Sr16 {830}. [Srrl2 {1238}]. 2B {830,1308}, 2BL {1307}. i: I Sr16-Ra {828}; I Th3B-Ra {832}. s: CS*7/Marquis 2B Sr9g {1581}; CS*4/Thatcher 2B Sr9g {1308}. CS*5/Thatcher 3B Sr12 {832}.

Sr17[{964}]. [sr17 {964}]. Recessive. 7B {771}, 7BL {964}. s: CS*6/Hope 7B {964}. v: Jing-Hong 1 Sr5 {564}; Suneca Sr2 Sr8a {485}. Lancer Sr2 Sr9d {679}; Scout Sr2 Sr9d {679}. Jing-Hong 2 Sr5 {564}. Nell Sr7b {1565}. E-Gan Zeo Sr8a {564}; Golden Valley Sr8a {979}. Present in many stocks possessing Pm5 {964}. See "Reaction to Erysiphe graminis.

Sr18 {054}. [SrG2 {844}, Srrl1 {1238}, Srmq1 {099}, SrPs1 {1263}, SrMn1 {1263}]. 1D {054,1308,1582}. i: I Hope 1D-Ra {828}, Sr18/8*LMPG {685}. s: CS*6/Hope 1D {1308}. v: Present in the majority of wheat stocks {828}.
Stocks not possessing Sr18: Brevit {054}; Chinese Spring {828}; Eureka {054}; Federation {054}; Gular {054}; Kenya C6042 {054}; Koala {054}; Little Club {828}; Morocco {054}; Norka {054}; Prelude {828}; Yalta {054}.
Sr19 {029}. [Srmq2 {099}]. 2B {029}, 2BS {1582}. v: Mq-B {029}. Marquis Sr7b Sr18 Sr20 {029}.

Sr20 {029}. [Srmq3 {1238}, Srrl3 {1238}]. 2B {029}. v: Mq-C {029}; Rl-C {029}. Reliance Sr5 Sr16 Sr18 {029}. Marquis Sr7b Sr18 Sr19 {029}.

Sr21 {1460}. 2AL {1460,1464}. i: Sr21/8*LMPG {685}. v: Hexaploid derivatives of T. monococcum {939}. tv: Tetraploid derivatives of T. monococcum {939}. dv: Einkorn {1460}. Various monococcum accessions. See also Sr45 which has similar specificity to Sr21.

Sr22 {1460}. 7A {649}, 7AL {1460}. i: Marquis*4//Stewart*3/T. monococcum {649,1460}; Sr22/9*LMPG {685}. Others {1112}. v: CS/3/Steinwedel*2//Spelmar/T. boeoticum {1460}; Schomburgk {880}; Steinwedel*2//Spelmar/T. boeoticum {1460}. Others {1112}. tv: Spelmar/T. boeoticum {1460}; Stewart*6/T. monococcum RL 5244 {649}. dv: Various T. monococcum accessions {649,1460}. ma: Hexaploid derivatives with Sr22 carried 'alien' segments of varying lengths; the shortest segment was distal to Xpsr129-7A {1112}.

Sr23 {950}. 4B {950}, 2BS {939}. These chromosome location results are consistant with the finding that the first location was based on Rescue monosomics. Rescue differs from CS by a 2B-4B reciprocal translocation {939}. v: Exchange {950}; Warden {950}. Etoile de Choisy Sr29 {950}. Sr23 is always associated with Lr16 {939}.

Sr24 {956}. Derived from Ag. elongatum.

3DL =T3DS.3DL-3Ae#1L {956,389}. i: Sr24/9*LMPG {685}; Sears' 3D/Ag translocations {956,1300}. v: Agent {956}; Blueboy II {956}; Collin {901}; Cloud {956}; Cody {1284}; Fox {956}; Gamka {785}; Karee {785}; Kinko {785}; Palmiet {785}; Sage {825,1024}; SST 23 {1324}; SST 25 {785}; SST 44=T4R {1324,785}; SST 102 {785}; Torres {128}; Wilga {785}. Siouxland Sr31 {1283}.

1BL.T1BL = 1BS-3Ae#1L] {600,389}. tr: Amigo {1463,600,389}; Teewon {600,389}. Note:Amigo also carries a 1AL.1RS translocation with resistance from rye {1463}.

3Ae#1. su: Chinese Spring 3Ag{3D} {1304}; TAP48 {389}.

Sr24 is completely linked in coupling with Lr24 {956} and often with red grain colour. See Reaction to P. recondita.

Sr25 {956}. Derived from Ag. elongatum.
7DL = T7DS.7DL-7Ae#1L {291,956,388,657}. i: Sears' CS 7D/7Ag translocations {956,1300};Sr25/9*LMPG {685}. v: Agatha Sr5 Sr9g Sr12 Sr16 {956} = T4 {1323}. Mutant 28 {388}. 7AL = T7A-7Ae#1L {330}. v: Sears' 7A/7Ae#1L No. 12 {330,1304}. Sears' 7D/7Ag#11 carries neither Sr25 nor Lr19 {939}. 7Ae#1L su: Chinese Spring + 7Ae#1L{7D} {1304}. See Lr19, reaction to Puccinia recondita.
Sr25/Lr19 often show complete linkage in wheat {956}.

Knott {681} obtained two mutants (28 and 235) of Agatha with reduced levels of yellow pigment in the flour. One of these mutants lacked Sr25. Marais {890} reported that a gene very similar to Sr25 was present in the putative Inia 66 x Thinopyrum distichum derivative, Indis. Marais {890,892} also obtained mutants with reduced yellow pigment in Indis derivatives and some of these lacked Sr25.

Sr26 {956}. Derived from Ag. elongatum. 6AL {364} = T6AS.6AL-6Ae#1L {388,389}. i: Sr26/9*LMPG{685}. v: Knott's 6A-6Ae#1L translocation to Thatcher {672}. Avocet {364}; Flinders {1449}; Harrier {939}; Jabiru {956}; King {1451}; Kite {956}; Takari {253}. Eagle Sr9g {956}. Bass Sr36 {1450}.

Sr27 Derived from S. cereale.

3A {T3A-3R} = T3AS.3R#1S {003,896,389}. i: Sr27/9*LMPG {685}. v: WRT wheat-rye translocation, available in CS, Thatcher and Pembina backgrounds. Translocated from Imperial rye to Chinese Spring by Acosta {003}. Widespread in triticales {966,1384}.
3B = T3BL.3R#1S {896}.

Sr28 {932}. 2BL {932}. i: Line AD {932}. v: Kota Sr7b Sr18 {932}.

Sr29 {313}. [SrEC {955}]. 6DL {313}, 6DS {1626}. i: Prelude/8*Marquis//Etoile de Choisy {313}. v: Hana {071}; Hela {076}; Mara {068}; Slavia {076}; Vala {076}. Etoile de Choisy Sr23 {955}.

Sr30 {688}. [SrW]. 5DL {688}. i: Sr30/7*LMPG - Lines 1, 2, and 3 {685}. v: Festiguay {688}; Mediterranean W1728 {1369}; Webster {688}. Klein Cometa Sr8b {1368}. Relatively common in Australian and Mexican wheats. Various unnamed accessions {208,1321}.

Sr31. Derived from S. cereale. 1B = T1BL.1RS = T1BL.& 1R#1S {389} or 1R{1B}. v: Derivatives of Petkus rye - see Lr26, Yr9. Feng-Kang 2 {563}; Feng-Kang 8 {563}; Gamtoos {785}; GR876 {753}; Jing-Dan 106 {563}; Lu-Mai 1 {563}; Jan 7770-4 {563}; Yi 78-4078 {563}. Amika {heterogeneous} Sr5 {076}; Dong Xie 3 Sr5 {563}; Dong Xie 4 Sr5 {563}; Istra Sr5 {076}; Solaris Sr5 {076}. Siouxland Sr24 {1283}. tv: Cando*2/Veery = KS91WGRC14 {381}.

Sr32. Dervied from T. speltoides.

2A {939,1304} = T2AL.2S#1L-2S#1S {389}. v: C95.24 {389}.
2B {1304} = T2BL-2S#1S {389}. v: C82.1 = P80-14.1-2 {389}.
2D {1304} = T2DL-2S#1L.2S#1S {389}. v: C82.2 = P80-139.1-4 {389,1304}; C82.3 = P80-132.2-2 {939,1304}; C82.4 = P80-153.1-2 {939,1304}.

Sr33. [SrSQ {650}]. 1DL {650}, 1DS {620} (linked with Gli-D1). v: RL 5405 = Tetra Canthatch/Aegilops squarrosa RL 5288 {650}.

Sr34 {967}. Derived from Ae. comosa.

2A {967} = T2AS-2M#1L.2M#1S {389}. v: CS 2A-2M 4/2 {967}.
2D {967} = T2DS-2M#1L.2M#1S {389}. i: Sr34/6*LMPG {685}. v: Compair {967}; CS 2D-2M 3/8 {967}. Various addition, substitution and translocation lines with Yr8 {967}.
2M {967}. su: Chinese Spring 2M(2A) {967}.

Sr35 {957}. [SrTm1 {1522}]. 3AL {957}. v,tv: Tetraploid and hexaploid derivatives of T. monococcum {957}. dv: T. monococcum C69. 69 Selection {957}; G2919 {957}.

Sr36 {939}. SrTt1 {949}. 2BS {939}. i: Sr36/8*LMPG {685}. v: T. timopheevii derivatives {949}. Arthur {939}; Arthur 71 {1324}; Flemink {1324}; Gouritz {1324}; Idaed 59; Maris Fundin {070}; Mengavi {949}; SST 101 {1324} SST 107 {785}; Timvera {949}; Zaragoza {785}. Timson Sr5 Sr6 {939}. Combination III Sr9e {939}. Bass Sr26 {1450}. Others {572}. tv: T. timopheevii {949}.

Sr37 {939}. [SrTt2 {949}]. 4BL {939}. v,tv: T. timopheevi and derivatives {484,949}. Line W {949}.

Sr38 {062}. Derived from Ae. ventricosa. 2AS {062}. i: RL 6081 =Thatcher + Lr37. This line will carry additional genes from Thatcher. v: VPM1.

Sr38 is linked with Lr37 and Yr17 - see Reaction to P. recondita and Reaction to P. striiformis

Sr39 {646}. Derived from Ae. speltoides. 2B {651}. v: RL 5711 {646,651}. tv: Amphiploid RL 5347 = Ae. speltoides/T. monococcum {651}.

Although Sr39 produces similar responses to Sr32, also derived from Ae. speltoides, recombination studies based on three crosses showed independent inheritance {646}. Sr39 segregated independently of Lr13 {651}.

Sr40 {302}. Derived from T. araraticum. 2BS {302} = T2BL/2G#2S {389}. i: RL 6087 = RL 6071*7/PGR 6126; RL 6088 = RL 6071*7/PGR 6195 {302}. tv: T. araraticum PGR 6126 {302}; PGR 6195 {302}.

Sr41 {1215}. 4D {1215}. v: WDR-B1 {1214}. Waldron Sr5 (heterogeneous) Sr11 (heterogeneous).

Sr42 {667}. 6DS {938}. v: Norin 40 {938}.

Sr43. Derived from Agropyron elongatum.

7DL = T7DL-7Ae#2L.7Ae#2S {657,389}. tr: KS10-2 {653}.
7D = T7DS.7Ae#2L {657,389}. tr: KS23-9 {653}; KS24-1 {653}; KS24-2 {653}.

Sr44 {389}. Derived from Agropyron intermedium.

T7DS-7Ai#1L.7Ai#S 7Ai#1L {389}. v: Line 86.187 {939}.
7Ai#2, &Ai#2S. su: Group 7 alien substitution lines with 7Ai#1 and 7Ai#1S {939}. ad: TAF2 = L1 {169}.

Sr45 {894}. SrD {934}; SrX {1805}. 1D {897}, 1DS {894}. v: 87M66-2-1 {894}; 87M66-5- 6 {897}; Thatcher + Lr21, RL5406 {894,934}. Various backcross derivatives developed at PBI Cobbitty {1461}. dv: T. tauschii RL5289 {894,934}.
Tests of natural and induced mutants of P. graminis f. sp. tritici indicated that Sr45 has identical specificity to Sr21 {934}.
Temporary designations:

SrA {323}. v: SW55-1 {323}; SW56-1 {323}. SW33-5 Sr9a Sr13 {323}. SW54-3 Sr9d Sr13 {323}.

SrTmp {1230}. v: Bai-Yu-Bao {564}; Beijing 9 {564}; Beijing 11 {564}; Fertodi 293 {977}; Martonvasari 5 {977}; Mironovskà = Mironovskaya 808 {068,977}; Nung-Ta 139 {564}; Parker {977}; Trison {1230}; Triumph 64 {841,1230,977}; Xuzhou 14 {564}; Yen-An 15 {564}. Beijing 10 Sr5 {564}.

SrWld {1230}. v: Prospect Sr11 {197}.

SrZdar {067}. 1B {067}. v: Zdar {067}.

Additional temporary designations are listed in {1230}.
Genotype lists: {323,970}.

Complex genotypes:
Centurk: Sr5 {979}, Sr6 {979}, Sr8a, Sr9a {979}, Sr17 {979}.
Chris: Sr5 {679,1371}, Sr7a {1371}, Sr9g {1371}, Sr12 {1371}.
Egret: Sr5 {939}, Sr8a {939}, Sr9b {939}, Sr12 {939}.
FKN: Sr2, Sr6, Sr7a, Sr8a {791}, Sr9b {791}.
H-44: Sr2, Sr7b {677}, Sr9d {677}, Sr17.
Hartog: Sr2 {127}, Sr8a, Sr9g, Sr12 {939}.
Hope: Sr2 {677}, Sr7b {677}, Sr9d {677}, Sr17.
Kenya Plume: Sr2 {1370}, Sr5 {1370}, Sr6 {1370}, Sr7a {1370}, Sr9b {1370}, Sr12 {1370} Sr17 {1370}.
Khapstein: Sr2, Sr7a, Sr13 {674}, Sr14 {674}.
Lawrence: Sr2, Sr7b {939}, Sr9d, Sr17.
Lerma Rojo 64: Sr2, Sr6, Sr7b {979}, Sr9a {979}.
Madden: Sr2, Sr9b, Sr11, Sr13 {842}.
Manitou: Sr5 {679}, Sr6 {679}, Sr7a, Sr9g {965}, Sr12 {939}.
Mendos: Sr7a {939}, Sr11 {879}, Sr17, Sr36.
PI 60599: Sr7a {689}, Sr8a, Sr9b, Sr10.
Selkirk: Sr2 {499}, Sr6 {468}, Sr7b {499}, Sr17, Sr23 {950}.
Redman: Sr2, Sr7b {939}, Sr9d {939}, Sr17.
Reliance: Sr5 {1308}, Sr16 {1238}, Sr18, Sr20.
Renown: Sr2, Sr7b {939}, Sr9d {939}, Sr17.
Roblin: Sr5 Sr7a? Sr11 Sr12
Timgalen: Sr5 {heterogeneous} {1555}, Sr6 {1555}, Sr8a, Sr36.
Thatcher: Sr5 {1308}, Sr9g {965}, Sr12 {939}, Sr16 {1308}.
WW15=Anza= Karamu=T4: Sr5 {939}, Sr8a {939}, Sr9b {939}, Sr12 {939}.

Reaction to Puccinia recondita Rob. ex Desm.
Disease: Brown rust, leaf rust.

Lr1 {047}. 1B {1409}, 5D {954}, 5DL {945}. i: Centenario/6*Thatcher {317}; Malakoff/6*Prelude {317}; Wichita*4/Malakoff {613}. v: Centenario {317}; Chicora 'S' {143}; Daws (heterogeneous) {1019}; Dirkwin {1019}; Glenlea {1255,976}; Halle 9H37 {074}; Hyslop {1019}; Luke {heterogeneous} {1019}; Malakoff {047}; McDermid {1019}; Mexico 120 {933}; Newton {143,1024,1023}; Norco {1019}; Shabati Sonora {842}; Sonora 64 {842}; Tarsa {842}; Uruguay {954}; Walliday {1019}. Erythrospermum 142 and 953 Lr3 {074}; Plainsman V Lr3 {1024}. Blueboy Lr10 {143}; Laura Lr10 Lr34 {712}. Suneca Lr13 {485}. Blueboy II Lr10 Lr24 {143}. Norka Lr20 {1552}. ma: Co-seg. with Xpsr567-5D and Xglk621-5D in a Frisal x Lr1 resistant line. pTAG621 was converted to a diagnostic STS {354}.

Lr2 1B {1409}, 2DS {843,942}.

Lr2a {320}. [Lr2 {047}]. i: Prelude*6/Webster {320}; Red Bobs*6/Webster {320}; Webster/6*Thatcher {306}; Wichita*4/Webster {613}. v: Eureka CI 17738 {143}; Festiguay {843}; Webster {047}. Mediterranean W1728 Lr3 {1369}. Shield Lr3 Lr10 {198}. Alex Lr10 {976}; Guard Lr10 {976}; James Lr10 {976}; Len Lr10 {976}; Marshall Lr10 {976}; Waldron Lr10 {143}.

Lr2b {320}. [Lr2² {1409}]. i: Prelude*6/Carina {320}; Red Bobs*6/Carina {320}; Thatcher*6/Carina {320}; Wichita/4*Carina {613}. v: Carina {613}.

Lr2c {320}. [Lr2³ {1409}]. i: Prelude*5/Brevit {320}; Prelude*6/Loros {320}; Red Bobs*6/Brevit {320}; Red Bobs*6/Loros {320}; Thatcher*4/Brevit {320}; Thatcher*6/Loros {320}; Wichita*4/Brevit {613}; Wichita*4/Loros {613}. v: Brevit {613}; Loros {317,1257}.

Lr3 {047}. 6B {549}, 6BL {939}. i: Democrat/6*Thatcher {318}; Wichita*4/ Mediterranean {613}. v: Belocerkovskaja 289 {074}; Bennett {1024}; Democrat {047}; Fertodi 293 {074}; Gage {1024}; Hana {068}; Homestead {1024}; Ilyitchovka {075}; Juna {075}; Jubilejnê {068}; Kawvale {143}; Lancota {1024}; Mara {068}; Mediterranean {047}; Mediterranean W3732 {1369}; Mentana {842}; Mironovskaya 264 & 808 {074}; Odra {075}; Osetinskaya {074}; Ottawa {143}; Pawnee {1408}; Ponca {143}; Rannaja 12 {074}; Shawnee {143}; Shirahada {842}; Skorospelka 3b {074}; Sledkovicova K1004 {074}; Viginta {068}; Warrior {143,1024}; Yubileynaya {075}. Erythrospermum 142 & 953 Lr1 {074}; Plainsman V Lr1 {1024}. Mediterranean W1728 Lr2a {1369}. Shield Lr2a Lr10 {198}. Bowie Lr14b {319}. Amika Lr26 {076}; Istra Lr26 {076}; Solaris Lr26 {076}. Bezostaya 1 Lr34 {074}; See also {069}.

Because Lr3 appears to be a complex locus {486} Democrat and Democrat/6*Thatcher should be accepted as standards. There is evidence to suggest that the allele in Mentana, and therefore many derivatives, is LrBg {939}.

Lr3bg{486}. i: Thatcher*6/Bage; RL6094 = Tc*6/T6 {307}. v: Bage {486}. T6 Lr16 {307}.

Lr3ka{486}. i: Tc*6/Klein Aniversario. v: Klein Aniversario {486}.

Lr4, Lr5, Lr6, Lr7, Lr8 {365}. v: Purdue Selection 3369-61-1-10 = Waban {365}. Not available as separate single-gene lines.

Lr9 {1408}. Derived from Ae. umbellulata. 6B {954,1296,1299}, 6BL = T6BS.6BL-6U#1L {389}. i: T47 = Transfer = CS + Lr9 {1408}; Thatcher*6/Transfer; Wichita*4/ Transfer {613}. v: Abe {143}; Arthur 71 {1320,1024}; Clemson 201 {465}; McNair 701 & 2203 {143}; PI 468940 {1439}; Riley 67 {1320,1024}; Sullivan {1110}. Oasis Lr11 {1109}. ma: Co-seg with XksuD27-6B {048}; co-seg with Xmwg684-6B and STS Xsfr1 {1272}; Lr9 - 8 cM - Xpsr546-6B {1272}.
The structures of additional translocations are given in {389}.

Lr10 {199}. [LrL {031}]. 1A {312,546}, 1AS {939}. i: Exchange/6*Thatcher {306}; Gabo/6*Thatcher {306}; Lee/6*Thatcher {306}; Selkirk/6*Thatcher {306}; Timstein/6*Thatcher {306}. s: CS*5/Timstein 1A {939}. v: Centurk {1024}; Centurk 78 {1024}; Concho {143}; Federation {939}; Kenya Farmer; Mayo 52 {031}; Mayo 54 {031}; Parker {546,1024}; Rocky {1024}; Sinton {1256}; Tascosa {143}; TAM-105 {055}. Blueboy Lr1 {143}. Blueboy II Lr1 Lr24 {143}. Laura Lr1 Lr34 {712}. Alex Lr2a {976}; Guard Lr2a {976}; James Lr2a {976}; Len Lr2a {976}; Marshall Lr2a {976}; Waldron Lr2a {143}. Shield Lr2a Lr3 {198}. Exchange Lr12 Lr16 {031}. Era Lr13 {143}. Selkirk Lr14a Lr16 {031,199}. Warden Lr16 {031}. Gabo Lr23 {031}; Lee Lr23 {031}; Timstein Lr23 {031}. Parker 76 Lr24 {143}; Unknown accessions {208}. ma: Xcdo426-1A - 5.1cM - Lr10 {1058}; Lr10 - 8cM - Glu-A3 {355}; cosegregation with Xsfr1(Lrk10)-1A and Xsfrp1(Lrk10)-1A {1270}; complete linkage with Lrk10, which encodes a protein kinase {639}.

Lr11 {1409}. 2A {1409}. i: Thatcher*6/Hussar {306}; Wichita*4/Hussar {613}. v: Bulgaria 88 {142}; Hart {1024}; Hazen {049}; Hussar {1409}. Oasis Lr9 {143}.

Lr12 {326}. Adult plant reaction. 4B {312}. i: Exchange/6*Thatcher {306}. v: Opal {306}. Sturdy Lr13 {301}. Chinese Spring Lr34 {301}. Exchange Lr10 Lr16 {326}. Unknown accessions {208}.

Lr13 {326}. 2BS {939}. Although originally described as a gene for adult plant reaction {032,326}, Lr13 can be detected at the seedling stage especially at high temperatures {939,1156}. i: Tc*7/Frontana = RL4031 {306}; fifteen Thatcher lines with 2-gene combinations {711}. v: This gene is very widespread {939}. Hustler {608}; Kinsman {608}; Kenya plume {1370}; Manitou {326}; Mardler {608}; Maris Huntsman {608}; Napayo {070}; Neepawa {143}; Polk {143}; Norman {608}; Virtue {608}. Yecora Lr1 {1374}. Hartog Lr1 {127}; Suneca Lr1 {485}. Klein Aniversario Lr3ka {032}. Era Lr10 {143}. Biggar Lr14a {712}; Genesis Lr14a {712}. Inia 66 Lr14a Lr17 {1373}. Columbus Lr16 {1258}; Kenyon Lr16 {300}. Hobbit Lr17 {608}; Lerma Rojo 64 Lr17 {1373}. Oasis 86 Lr19 {1373}. Cumpas 88 Lr26 {1373}. Chris Lr34; Frontana Lr34 {032,326,1374}; Parula Lr34 {1374}.

Lr14.

Lr14a {319,964}. 7B {964}, 7BL {770}. i: Selkirk/6*Thatcher {319}. s: CS*6/Hope 7B {964}. v: Aotea {964}; Brigand {608}; Gala {964}; Glenwari {964}; Hofed {964}; Hope {964}; H-44 {964}; Lawrence {964}; Redman {964}; Regent {964}; Renown {964}; Spica {964}. Selkirk Lr10 Lr16 {319}. Biggar Lr13 {712}; Genesis Lr13 {712}. Inia 66 Lr13 Lr17 {939}.

Lr14b {319}. i: Maria Escobar/6*Thatcher {319}. v: Bowie Lr3 9226}. Maria Escobar Lr17 {319}; Rafaela Lr17 {314}.

Lr14ab i: Lr14a/6*Thatcher//Lr14b/6*Thatcher {319}.

Lr15 {843}. 2DS {843,942}. i: Thatcher*6/Kenya W1483 {306}. v: Kenya W1483 {843}.

Lr16 {318}. [LrE {031}]. 4B {312}, 2BS {939}. Note: These chromosome locations are consistant with the finding that the first location was based on the use of a Rescue monosomic series. Rescue differs from CS by a 2B-4B translocation {939}. i: Exchange/6*Thatcher {306}; RL6096 = Tc*6/T6 {307}. v: Ciano 79 {1373}. Etoile de Choiosy {074}; Imuris 79 {1373}; Papago 86 {1373}. T6 Lr3bg {307}. Warden Lr10 {031}. Exchange Lr10 Lr12 {031}. Selkirk Lr10 Lr14a {031}. Columbus (heterogeneous) Lr13 {1258}; Kenyon Lr13 {300}.

Lr16 is always asociated with Sr23.

Lr17 {318}.

Lr17a [{318}], {1350}. [Lr17]. 2A {314}, 2AS {062}. i: Klein Lucero/6*Prelude {318}; Klein Lucero/6*Thatcher {318}; Maria Escobar/4*Thatcher {318}. v: EAP 26127 {314}; Jupateco {939}; Klein Lucero {318}. Hobbit Sib Lr13 {939}; Lerma Rojo 64 Lr13 {1373}. Inia 66 Lr13 Lr14a {9010}. Maria Escobar Lr14b {318}; Rafaela Lr14b {314}.

Lr17b {1350}. [LrH {970}, WBR2 {615}]. v: Harrier {1350}; Norin 10-Brevor, 14 {1350}; Maris Fundin {1350}. Hobbit Sib = Dwarf A Lr13 {1350}.

Lr18 {318}. 5BL {935} = T5BS.5BL-5G#1L {389}. i: Africa 43/7*Thatcher {318}; Red Egyptian PI 170925/6*Thatcher {318}. v: Africa 43 {318}; Red Egyptian PI 170925 {318}; Red Egyptian PI 17016-2c {318}; Sabikei 12 {935}; Timvera {935}; Timvera Derivative. {935}. Certain WYR accessions {935}. FTF {1614}; Several Sabikei lines including Sabikei 12 {1614}.

Independently derived lines with Lr18 possess a unique N band terminally located in chromosome 5BL {1614}. Low seedling responses conferred by Lr18 are most effective at 15-18oC. With increasing temperatures the response becomes less effective and ineffective at 25-27oC {935, see also, 1614}.

Lr19 {140}. Derived from Ag. elongatum.

7DL = T7DS.7DL-7Ae#1L {291,956,1323,388,657,389}. i: Agatha = T4 = TC + Lr19 {956,1323}. v: L503 {1346}; L513 {1346}; Mutant 28 {598}. Sunnan {684}. Oasis 86 Lr13 {1373}. See Sr25. ma: Co-seg with 8 RFLP markers {048}; Ep-D1c - 0.33 cM - Lr19 {1587}; cosegregation with Ep-D1d {974}. Prins et al. {1162} studied 29 deletion mutants in Indis and determined the gene order: Sd-1 - Xpsr105 - Xpsr129 - Lr19 - Wsp- D1 - Sr25 - Y.
7DL = T7DS.7DL-7Ae#1-7DL {388}. v: Mutant 235 {681};
7AL = T7A-7Ae#1 {330}. v: Sears' 7A-7Ag No. 12 {330}.
7BL {1163} v: 88M22-149 {1163}.
7AgL {1304} = 7Ae#1L. su: Agrus.
Lr19 is usually associated with Sr25. However Sear's transfer 7D-7Ag No. 11 carries neither Lr19 nor Sr25. SeeLr29.

Knott {681} obtained two mutants (28 and 235) of Agatha possessing Lr19, but with reduced levels of yellow pigment in the flour. Marais {890,892} obtained mutants and recombined lines with intermediate levels of, or no, yellow pigment. It was shown that in recombinant line 88M22-149 lacking yellow pigment, Lr19 was transferred to chromosome 7BL {1163}.
The chromosome with Lr19 in Indis is probably identical to that in Agatha {1162}.

Lr20 {140}. 7AL {1305,1554}. s: CS*5/Axminster 7A {1293}. v: Axminster {348,1175,1305}; Birdproof {1554}; Bonus {1554}; Converse {1554}; Festival {1554}; Kenora {1554}; Kenya W744 {1554}; Maris Halberd {608}; Normandie {348,1554}; Sappo {608}; Sicco {310}; Thew {140,1552}; Timmo {608}. Norka Lr1 {1554}. See Pm1 (Reaction to Erysiphe graminis) & Sr15 (Reaction to Puccinia graminis) with which Lr20 is always associated. Lr20 in Sicco appears to differ from that in Sappo, Timmo and Maris Halberd {310}. Lr20 in Norka (Lr1+Lr20) may differ from that in thew {939}.

Lr21 {1241}. 1D {650}, 1DL {1241}, 1DS {448}. i: Thatcher*6/Tetra Canthatch/Ae. squarrosa var. meyeri RL 5289 {306}. v: Tetra Canthatch/Ae. squarrosa var. meyeri RL 5289, RL 5406 {648}. AC Cora Lr13 {713}.

Lr22 2DS {1241}.

Lr22a {1241}. Adult plant reaction. i: Thatcher*3//Tetra Canthatch/Ae. squarrosa var. strangulata RL 5271 {306}. v: Tetra Canthatch/Ae. squarrosa var. strangulata RL 5271, RL 5404 {311}. AC Minto Lr11 Lr13 {713}. dv: Ae. squarrosa var. strongulata RL 5271.

Lr22b {298}. Adult plant reaction. v: Canthatch {298}; Marquis {970}; Thatcher {298}. This gene will be present in near-isogenic lines based on Thatcher.

Lr23 {948}. [LrG {951}]. 2BS {948}. i: Lee FL 310/6*Thatcher {948}. s: CS*7/Kenya Farmer 2B {948}; CS*6/Timstein 2B {948}. v: Crim {1091}; Hope/Timstein {1091}; I 310678 {1091}; I 310685 {1091}; I 349162 {1091}; K 45973 {1091}; K 51070 {1091}; Rocta {1091}. Gamenya Lr3 {1552}. Gabo Lr10 {1552}; Kenya Farmer Lr10 {1552}; Lee Lr10 {1552}; Timstein Lr10 {1552}. tv: Altar 84{1058}.

Suppressor of Lr23:

SuLr23 {1058}. Suppressing allele. 2DS {1058}. v: Altar 84/T. tauschii 219 {1058}.

suLr23 {1058}. v: Opata 85 {1058}.

Lr24. [LrAg {141}]. Derived from Ag. elongatum.

Always present with Sr24 {956}. See Sr24(Reaction to Puccinia graminis).

3D {956,1389}. v: Cody {1284}; Osage {143}; Payne {1390,1024}; SST 23 {1324}; SST 44 = T4R {1324}; Timpaw {1255}; Torres {128}; Wanken {1255}. Blueboy II Lr1 Lr10 {141}. Fox Lr10 {141}; Parker 76 Lr10 {143,1024}. Siouxland Lr26 {1283}. ma: Co-seg of Lr24 in Agent with 8 RFLP markers; segment in Sears’ 3D-3Ag#1 is shorter than in Agent {048}. ma: Tagged with Xpsr1203-6B {1271}; cosegregation with RAPD marker that was converted to a SCAR {231}.
1BL {185} = T1BL.1BS-3Ae#1L {600}. v: Amigo {1463,600,185}; Teewon {600}.
Lr25. Derived from S. cereale cv. Rosen.

4BS {270,271,380,389}; revised to T4BS.4BL-5RL {543} and later to T4BS.4BL-2R#1L. v: Transec {273}; Transfed {269}. Always present with Pm7. ma: Cosegregation with a RAPD {1165}.

Lr26. 1R{1B}, T1BL-1RS. v: Derivatives of Petkus rye - see Yr9 (Reaction to Puccinia striifromis) & Sr31 (Reacton to Puccinia graminis). Iris {075}; Sabina {075}; GR876 {753}; Bacanora 88 {1373}. Many wheats with Lr26 also carry Lr3. Amika {heterogeneous} Lr3 {076}; Istra Lr3 {076}; Solaris Lr3 {076}. Cumpas 88 Lr13 {1373}. Siouxland Lr24 {1283}. See also {310}. tv: Cando*2/Veery, KS91WGRC14 {381}.

Hanusova et al. {492} identified 127 wheats with Lr26 but only 16 of them were listed.

Lr27 {1367}. [LrGt, A {1366}]. 3BS {1367}. One of two complementary genes; the second gene, Lr31, is located in chromosome 4BS {1367}. The following wheats have both Lr27 and Lr31. s: CS*6/Ciano 3B {1366}; CS*6/Ciano 5B {1366}; CS*6/Hope 3B {1366}. v: Gatcher {1366}. Ocoroni 86 {1373}. SUN 27A Lr1 Lr2a {1366}; Timgalen Lr3 {heterogeneous} Lr10 {1366}. Anhuac Lr13 Lr17 {1361}; Cocoraque 75 Lr13 Lr17 Lr34 {1361}; Jupateco 73S Lr17 {1361}.

Lr27 is present in wheats with Sr2, but is not expressed in the absence of the complementary factor {1366}. ma: Positive association with XksuG53-3B {1058}.

Lr28 {967}. Derived from Ae. speltoides.

4AL {967} = T4AS.4AL-7S#2S{389}. i: CS 2A/2M 4/2 {967}; CS 2D/2M 3/8 {967}. ma: Lr28 was tagged using STS primer OPJ-02378 {1052}.

Lr29 {939}. Derived from Ag. elongatum.

7DS {939} = T7DL-7Ae#1S {389}. i: Sears' CS 7D/Ag#11 {939,1300}; RL6080 = Tc*6/Sears’ 7D/Ag#11 {316}. ma: Cosegregation with two RAPDs {1165}.

Lr30 {315}. [LrT]. Recessive {315}. 4AL {315}. i: RL 6049 = Thatcher*6/Terenzio, {315}. v: Terenzio {315}.

Lr31 {1367}. [B {1366}]. 4BL {1367}. One of two complementary genes, the second gene is Lr27. v: Chinese Spring {1366}. Ocoroni 86 {1373}. See Lr27 for list of wheats with Lr27 +Lr31.
ma: A positive association with XksuG10-4B {1058}.

Lr32. 3D {644}, 3DS{645}. v: Tetra Canthatch/T. tauschii RL 5497-1; RL 5713, RL 5713/Marquis-K {644}. dv: T. tauschii RL5497-1 {644}. ma: Xbcd1278-3D - 3.6 cM - Lr32 {048}; Xcdo395-3D - 6.9 cM - Lr32 {048}.

Lr33 {325}. 1BL {325}. i: RL 6057 = Tc*6/PI 58548 {297,325,321}. v: PI 268454a {297}; PI 58548 {297,325}. PI 268316 Lr2c Lr34 {297}. Others {1322}.

Lr34 {297,299}. [LrT2 {321}]. 7D {299}, 7DS {324,1058}. i: Line 897 {321}; Line 920 {321}. RL 6058 = Tc*6/PI 58548 {297}. RL 6059 = Tc*6/Terenzio Lr33 {297}. RL 6069 = Tc*6/Lageadinho LrT3 {321}; RL 6070 = Tc*5/PI 321999 LrT3 {321}; RL 6050 = Tc*6/Terenzio LrT3 {321}. Selections Jupeteco 73R Lr17 Lr27 + Lr31 and Jupateco 73S Lr17 Lr27 + Lr31 and Cocoraque 75 Lr13 Lr17 Lr27 + Lr31 and Anhuac 75 Lr13 Lr17 Lr27 + Lr31, can be considered near-isogenic for the presence and absence respectively, of Lr34 {1361}. Thirteen Thatcher lines with 2-gene combinations {434}. v: PI 268454 {297}. Glenlea Lr1 {327}. Laura Lr1 Lr10 {712}. Terenzio Lr3 Lr30 LrT3 {321}. Chinese Spring Lr12 {301}. Sturdy Lr12 Lr13 {301}. Frontana Lr13 {1374}; Parula Lr13 {1374}. PI 58548 Lr33 {297,321}. Lageadinho LrT3 {321}. Others {299,321,1322,1376}. See {1362}. ma: Complete linkage with Ltn {1361} (leaf tip necrosis), Yr18 (Reaction to Puccinia striiformis) {1362,937} and Bdv1 (Reaction to barley yellow dwarf virus).

In addition to conferring seedling and adult plant resistance, Lr34 responds in a complementary manner when combined with either Lr33 or LrT3 {321}. In the Thatcher background, Lr34 is associated with increased resistance to stem rust {299,321}.

Although the resistance gene in the near-isogenic Thatcher line, RL6077, was considered to be Lr34 on the basis of disease response, leaf tip necrosis and its association with resistance to stripe rust, a cross with RL6058 segregated for two genes. A translocation to another chromosome was suggested {324}.

Lr35 {651}. 2B {651}. v: RL 5711 {651}.

Lr36. Derived from Ae. speltoides. 6BS {292}. v: Line 2-9-2; Line E84018. al: Ae. speltoides Popn. 2.

Lr37 {062}. Derived from T. ventricosum. 2AS {062}. i: RL 6081 = Tc*/VPM1 {939}. RL6081 = Tc*8/VPM1 {316}. v: Hyak {021}; Madsen {020}; Rendezvous {062}; VPM1 {062}; VPM1 derivatives {939}.

Lr37 can be recognised in seedlings at low temperatures (17C) and is effective in adult plants under field conditions. See also Sr38 (Reaction to Puccinia graminis) and Yr17 (Reacton to Puccinia recondita)

Lr38 {392}. Derived from Ag. intermedium.

1DL = T1DS.1DL-7Ai#2L {390,389}. v: T25 {390}.
2AL = 2AS.2AL-7Ai#2L {392,389}. v: W49 {392} = T33 {390}.
3DS = 3DL.3DS-7Ai#2L {390,389}. v: T4 {390}.
5AS = 5AL.5AS-7Ai#2L {390,389}. v: T24 {390}.
6DL = 6DS.6DL-7Ai#2L {390,389}. i: RL6097 = Thatcher*6/T7 {307}. v: T7 {390,307}. 7Ai#2(7D) {392,389}. 7Ai#2(7A) {390}. su: W52 {390,389}.

Lr39 {1200}. Proposal under discussion.

Lr40 {1200}. Proposal under discussion.

Lr41 {215}. 1D {215}. v: KS90WGRC10 = TAM107*3/T. tauschii TA2460 {220}. dv: TA 2460 {220}.

Lr42 {218}. 1D {218}. v: KS91WGRC11 = Century*3/T. tauschii TA2450. dv: T. tauschii TA2450.

Lr43 {218}. 7D {576}, 7DS {577}. v: KS92WGRC16 = Triumph 64/3/KS8010-71/TA2470//TAM200. dv: T. tauschii TA2470.

Lr44 {322}. 1B {322}. i: RL = 6147 Thatcher*6/T. spelta 7831 {322}. v: T. spelta 7831 {322}; T. spelta 7839 {322}.

Lr45 {958}. Derived from Secale cereale. 2A = T2AS-2R#3S.2R#3L {958,389}. i: RL6144 = Thatcher*7/ST-1 {958}. v: ST-1 {958}. Various Australian backcross derivatives {958}.

Lr46 {1364}. 1B {1364}. s: Lalbahadur(Pavon 1B) Lr1 {1364}. v: Pavon F76 Lr1 Lr10 Lr13 {1364}.

Temporary designations:
LrTb {820}. Adult plant resistance {820}. v: AC Taber Lr13 Lr14a {820}.

LrVPM {1603}. 7DL {7411}.

LrW {309}. v: V618 Lr33 {309}. V336 Lr33 LrB {309}.

LrW2 {305}.
A gene, identified only as Lr, was transferred to wheat chromosome 2AS from 6Mv {113}. Cosegregating markers were Xpsr933-2A and Xpsr150-2A.

A series of temporary designations for seedling and adult plant resistance genes in six durums is given in {1648}.
Complex genotypes:

AC Domain: Lr10 Lr16 Lr34 {820}.
Benito: Lr1 Lr2a Lr12 Lr13 {1256}.
Buck Manantial: Lr3 Lr13 Lr16 Lr17 Lr34? {300}.
Era: Lr10 Lr13 Lr34 {342}.
Grandin: Lr2a Lr3 Lr10 Lr13 Lr34 {821}.
Mango: Lr1 Lr13 Lr26 Lr34 {1374}.
MN7529: Lr1 Lr2a Lr10 Lr16 {976}.
Opata 85: Lr10 Lr27+Lr31 Lr34 {1058}.
Pasqua: Lr11 Lr13 Lr14b Lr30 Lr34 {304}.
Prospect: Lr1 Lr2a Lr10 Lr13 {197}.
Roblin: Lr1 Lr10 Lr13 Lr34 {303,713}.
Trap: Lr1 Lr3 Lr10 Lr13 Lr34 {1374}.

Genotype lists: {1361} (combinations with Lr34), {855} (Czechoslovakian cultivars), {1365} (Indian Subcontinent), {1219,978}, (U.S.A. cultivars), {1365,1345} (Indian cultivars), {1373} (Mexican cultivars), {1380} (cultivars from the former USSR), see alse {970}.

Reaction to Puccinia striiformis Westend.

Disease: Stripe rust, yellow rust.

Yr1 {851}. [L {1622}]. 2A {877,1610}, 2AL {940}. v: Chinese 166 {851}; Corin {230}; Dalee {083}; Durin {1459}; E2025 {1267}; E7700 {1267}; E8594 {1267}; Feng-Kang 13 {1610}; Heines 110 {604}; Maris Ranger {1459}; Maris Templar {1459}; Odra {073}. Fenman Yr2 {1459}; Ibis Yr2 {604}; Rothwell Perdix Yr2 {604}; Regina Yr2 {073}; Sylvia Yr2 {1430}; Tadorna Yr2 {1431}. Avocet {UK} Yr2 Yr6 {1459}; Longbow Yr2 Yr6 {083}; Mithras Yr2 Yr6 {1459}. Marksman {heterogeneous} Yr2 Yr13 {1459}. Galahad Yr2 {heterogeneous} Yr14 {1459}. Mardler Yr2 Yr3a Yr4a Yr13 {604,1459}. Nudif TP3 Yr3c {1431}. Nudif TP1 Yr3a {1431}. Maris Templar Yr3a Yr4a {604}. Argent Yr3a Yr4a Yr6 {1067}. Nudif TP250 Yr6 {1431}. Stetson Yr9 {083}. Bounty Yr13 {1459}; Sportsman Yr13 {1459}; Virtue Yr13 {083,1459}. Galahad Yr14 {083}.

A report {1267} that Kalyansona and Nadadores carried Yr1 is not correct.

Yr2 {851}. [U {1622}]. Recessive {1351}. 7B {746,186,184}. v: Derius {230}; Flevina {1431}; Hana {51,58}; HD2329 {1352}; Heines VII {851}; Kalyansona {1352}; Laketch {050}; Leda {1430}; Manella {1431}; Merlin {1622}; Odra {071}; PBW54 {1352}; PBW120 {1352}; Slavia {073,071}; Soissonais Desprez {851}; WG377 {1352}; WH147 {1352}; WL711 {1352}; WL1562 {1352}; Yamhill {181}. Fenman Yr1 {1459}; Ibis Yr1 {604}; Rothwell Perdix Yr1 {604}; Sylvia Yr1 {1430}; Tadorna Yr1 {1431}. Mardler Yr1 Yr3a Yr4a Yr13 {1459}; Mithras Yr1 Yr6 {1459}. Avocet (U.K.) Yr1 Yr6 {1459}; Longbow Yr1 Yr6 {083}; Marksman Yr1 {heterogeneous} Yr13 {1459}. Galahad (heterogeneous) Yr1 Yr14 {1459}; Viginta Yr3a Yr4a {073,071}; Maris Huntsman Yr3a Yr4a Yr13 {604}. Maris Beacon Yr3b Yr4b {1459}. Cleo Yr3c Yr14 {1431}. Yamhill Yr3a Yr4a {181,182, see also, 184}. Cleo Yr3c {1457}; Liberator Yr3c {1431}. Stella Yr3 {1430}. Rapier Yr4 {083}. Zdar Yr4a {073}. Flamingo Yr6 {1430}; Heines Kolben Yr6 {611}; Heines Peko Yr6 {746}; Norman Yr6 {083}. Flevina Yr7 {1430}; Garant Yr7 {230}; Hardi Yr7 {230}; Lely Yr7 {1430}. Nautica Yr9 {1430}. Maris Nimrod Yr13 {1459}. Brigand Yr14 {083}; Wizard {heterogeneous} Yr14 {1459}. Sonalika YrA {1352}.
Yr2 originally referred to a gene in Heines VII conferring resistance to European pathotypes. However, Heines VII possesses an additional resistance gene Yr25 {1351} which can be detected with a geographically wider range of pathogen isolates. Yr2 is present in Kalyansona {1351} and a range of spring wheats distributed by CIMMYT.

Yr3.

Yr3. Undesignated allele. v: Enkoy {050}. Vilmorin 23; Staring {1430}. Stella Yr2 {1430}; Minister Yr2 {1430}.

Yr3a {851}. 1B {185,184}. v: Bon Fermier {1431}; Nudif TP1 {1431}. Mardler Yr1 Yr2 Yr4a Yr13 {1459}. Argent Yr1 Yr4a Yr6 {1067}. Viginta Yr2 Yr4a. Yamhill Yr2 Yr4a {182}. Maris Huntsman Yr2 Yr4a Yr13 {604}. Cappelle-Desprez Yr4a {851}. Nord Desprez Yr4a {182,184}; Stephens {182,184}; Druchamp Yr4a {185,182, see also, 184}; Top Yr4a {230}. Zdar Yr4a {073,071}. Kinsman Yr4a Yr6 {604}; Maris Freeman Yr4a Yr6 {604}; Maris Ranger Yr4a Yr6 {604}. Hobbit Yr4a Yr14 {604}.
Yr3b {851}. v: Hybrid 46 Yr4b {851}. Chen & Line {182} found that a second gene in Hybrid 46 - presumably this gene - was not located at the Yr3 locus.
Yr3c {851}. 1B {184}. v: Minister {851,182,184}. Cleo Yr2 {1430}. Maris Beacon Yr2 Yr4b {1459}.

Yr4

Yr4. Undesignated allele. v: Kenya Kubangu {050}. Rapier Yr2 Yr14 {083}. Avalon Yr14 {083}.

Yr4a {851}. 6B {185,184}. v: Vilmorin23 {184}. Mardler Yr1 Yr2 Yr3a Yr13 {1459}. Argent Yr1 Yr3a Yr6 {1067}. Yamhill Yr2 Yr3a {182,185, see also, 184}. Viginta Yr2 Yr3a {073,071}. Huntsman Yr2 Yr3a Yr13 {604}. Cappelle-Desprez Yr3a {851}; Druchamp Yr3a {182}; Nord Desprez Yr3a {182}; Top Yr3a {230}; Maris Freeman Yr3a Yr6 {604}; Kinsman Yr3a Yr6 {604}; Maris Ranger Yr3a Yr6 {604}. Hobbit Yr3a Yr14 {604}. Zdar Yr3c {073,071}.

Yr4b {851}. 6B {184}. v: Avalon {1160}; Opal {1431}; Staring {1430}; Stella Yr2 {1430}. Maris Beacon Yr2 Yr3b {1160,1459}. Hybrid 46 Yr3b {851,182, see also, 184}. Nudif TP12 Yr3c {1431}.

Yr5 {877}. 2BL {034}. v: E5557 {1267}; E8510 {1267}; T. spelta album {877}. Seven spelt accessions from Europe and Iran {640}.

Yr6 {877}. [B {1622}]. 7B {746}, 7BS {331}. v: Austerlitz {230}; Fielder {181}; Heines Kolben {1622}; Koga II {746}; Maris Dove {604}; Récital {230}; Takari {368}. Nudif TP250 Yr1 {1431}. Avocet (UK) Yr1 Yr2 {1459}; Longbow Yr1 Yr2 {083,1459}; Mithras Yr1 Yr2 {1459}. Argent Yr1 Yr3a Yr4a {1067}. Heines Peko Yr2 {746,877}; Flamingo Yr2 {1430}; Kolben Yr2 {611}; Norman Yr2 {083,1459}. Kinsman Yr3a Yr4a {604}; Maris Freeman Yr3a Yr4a {604}; Maris Ranger Yr3a Yr4a {604}. Orca Yr3c {1431}. Nudif TP241 Yr7 {1431}; Pavon 76 Yr7 {284}. Penjamo 62 (heterogeneous) Yr34 {1562}. tv: Duilio {192}; Latino {192}; Norba {192}; Quadruro {192}; Rodeo (heterogeneous) {192}.

Yr7 {877}. 2B {612,1429}, 2BL {965}. v: Present in many hexaploid wheats with Sr9g - see {965}. Brock {083}; Lee; Nudif TP257 {1431}; PBW12 {1352}; Prinqual {230}; Renard {083}; Talent {230}; Tango {230}; Tommy {083}; WL2265 {1352}. Flevina Yr2 {1431}; Garant Yr2 {230}; Hardi Yr2 {230}; Lely Yr2 {1430}. Nudif TP241 Yr6 {1431}; Pavon 76 Yr6 {284}. Donata Yr9 {1430}; Pakistan 81 = Veery#5 Yr9 {284}. tv: Iumillo {965}, but not present Acme and Kubanka which also carry Sr9g {965}.

Yr8 {1217,1218}. Derived from Ae. comosa.

2D = T2D-2M {1218} = T2DS-2M#1L.2M#1S {389}. tr: Chromosome 2D-2M translocations in Hobbit Sib and Maris Widgeon {1016}. Compair {1217,1218}; CS 3D/2M 3/8 {967}. See also Sr34 and {967}.
2A = 2A-2M = T2AS-2M#1L.2M#1S {389}. tr: CA 2A/2M 4/2 {967}.
2M-1. su: CS 2M#1(2A) {967}.

Yr9 {878}. Derived from S. cereale.

1B = 1RS.1BL. v: Almus {998}; Aurora {1623}; Benno {998}; Bezostaya II {998}; Clement {1430,1532}; GR876 {753}; Kavkaz {1623}; Lovrin 10 {998}; Lovrin 13 {998}; Mildress {1027}; Perseus {998}; Predgornaya {998}; Salmon {998}; Sarhad 82 {284}; Skorospelka 35 {998}; Veery {986}; Weique {1627}; Winnetou {998}; Riebesel 47/51 {878,1623}; St 2153/63 {997}; Weihenstephan 1007/53 {1623}. Pakistan 81 = Veery#5 Yr7 {284}. tv: Cando*2/Veery, KS91WGRC14 {381}.

1R(1B) {997,1623}. su: Burgas 2 {998}; Clement {1160}; Lovrin 13 {998}; Mildress {998}; Mironovskaja 10 {998}; Neuzücht {1623}; Orlando {1623}; Roseana {068}; Saladin {997}; Salzmünder Bartweizen {1623}; St 14/44 {998}; Weique {1627}; Wentzel W{1623}; Winnetou {1027}; Zorba {998}. See also {050}.

Chromosome status not specified. v: Baron {083}; Branka {071}; Danubia {068}; Hammer {083}. Iris {068}; Kromerzhizhskaya {1149}; Lyutestsens 15 {1149}; Roxana {068}; Sabina {068}. Selekta {068}; Shtorm {1149}. Solaris {068}; Stuart {083}. Stetson Yr1 {083}. Nautica Yr2 {1430}; Agra Yr3 {068,071}; Sparta Yr3 {071}. Donata Yr7 {1430}.

Yr10 {878}. 1B {641}, 1BS {1002}. v: Moro {878}; PI 178383 {878}; T. spelta 415 {641}.

Yr11. [R11 {1157}]. Adult plant resistance. v: Joss Cambier {606}.

Yr12. R12 {1157}. Adult plant resistance. v: Fleurus {1158}; Frontier {1159}; Pride {1157}. Armada Yr3a Yr4a {1160,081}; Mega Yr3a Yr4a {1157,1160}. Waggoner Yr3a Yr4a Yr6 {1158}.

Yr13. R13 {1157}. Adult plant resistance. v: Hustler Yr1 Yr2 Yr3a Yr4a {083,1459}; Mardler Yr1 Yr2 Yr3a Yr4a {1459}; Marksman Yr1 {heterogeneous} Yr2 Yr3a Yr4a {1459}. Bounty Yr1 Yr3a Yr4a {1459}; Sportsman Yr1 Yr3a Yr4a {1459}; Virtue Yr1 Yr3a Yr4a {083,1158,1459}. Guardian Yr2 {082}. Maris Huntsman Yr2 Yr3a Yr4a {083,604,1459}; Pageant Yr2 Yr3a Yr4a {082}; Professor Marchal Yr2 Yr3a Yr4a {607}; Maris Nimrod Yr2 Yr3a Yr4a {607,1157,1459}. Brigand Yr2 Yr3a Yr4a Yr14 {609}; Gawain Yr2 Yr3a Yr4a Yr14 {081}. Copain Yr3a Yr4a {1158}; Kinsman Yr3a Yr4a Yr6 {1459}.

Yr14. [R14 {1157}]. Adult plant resistance. v: Kador {1158}; Score {1157}; Wembley {610}. Galahad Yr1 Yr2 (heterogeneous) Yr3a Yr4a {083,1459}. Brigand Yr2 Yr3a Yr4a Yr13 {083,609,1459}; Gawain Yr2 Yr3a Yr4a Yr13 {081}. Rapier Yr2 Yr3b Yr4b {083}; Wizard Yr2  (heterogeneous) Yr3b Yr4b {083,1459}. Hobbit Yr3a Yr4a {1459,1157}; Maris Bilbo Yr3a Yr4a {1157,1459}. Avalon Yr3b Yr4b {083,1459}. Moulin Yr6 {083}.

Yr15 {432,969}. 1BS {939,969}. v: Hexaploid derivatives of T. dicoccoides G-25 {432,466}; V763-2312 {969}; V763-254 {969}. tv: T. dicoccoides G-25 {432,431,466}; D447 derivatives B1, B2, B9, B10 {1434}. ma:tv: OPB131420 - 27.1 cM - Yr15 - 11.0 cM - Nor-B1 {1434}.

Yr16 {1598}. Adult plant resistance. 2D {1598}. v: Bersee {1604}; Cappelle-Desprez {1598}.

Yr17 {062}. 2AS {062}. v: See Lr37 (Reaction to Puccinia recondita) and Sr38 (Reaction to Puccina graminis). RL 6081 probably carries Yr7.

Yr18 {1362}. 7D {1362}; 7DS {324}. i: Thatcher near-isogenic lines with Lr34 including the 13 2-gene combinations reported in {434,937}. v: Jupateco 73R; Lerma Rojo 64 {1375}; Nacazari 76 {1375}; Tesia F 79 {1375}; Tonichi S 81 {1375}; Wheaton {1375}. Penjamo 62 Yr6 (heterogeneous) {1375}. Wheats with Lr34 (See Lr34). Others {1376}. ma: Complete linkage with Lr34 {937,1362}, Ltn {1361} and Bdv1 {1363}.

Yr19 {183}. [YrCom {183}]. 5B {183}. v: Compair Yr8 {183}.

Yr20 {183}. [YrFie {181}]. 6D {183}. v: Fielder Yr6 {183}.

Yr21 {183}. [YrLem {181}]. 1B {183}. v: Lemhi {183}.

Yr22 {183}. [YrLe1 {183}]. 4D {183}. v: Lee Yr7 Yr23 {183}.

Yr23 {183}. [YrLe2 {183}]. 6D {183}. v: Lee Yr7 Yr22 {183}.

Yr24 {952}. 1BS {952}. v: Meering*3/K733/T. tauschii AUS18911 {952}. tv: K733 {952}.

Yr25 {158}. 1D {158}. v: TP1295 {158}; TP981{158}.

Although this gene is likely to be present in Strubes Dickkopf, Heines VII Yr2, Heines Peko Yr2 Yr6, Reichersberg 42 Yr7 and Clement Yr9 {158}, the foregoing lists were not adjusted accordingly.

Yr26 {617}. 6AS (6AL.6VS) {617}. Derived from Haynaldia villosa (Daspyrum villosum). v: Yangmai-5 {617}.

Yr27 {953}. [YrSk {928}]. 2BS {953}. v: Ciano 79 {953}; Selkirk {953}.
Yr27 is present in many CIMMYT wheat lines {953} and possibly Webster. Note: Yr27 is closely linked with Lr13 (repulsion).

Yr28 {1377}. 4DS {1377}. v: Synthetic = Altar 84/T. tauschii W-219. Synthetic/Opata 85 SSD. dv: T. tauschii W-219 {1377}.
Yr22 was also reported for chromosome 4D but in the absence of an appropriate single gene stock and the unavailability of avirulent cultures in most laboratories, tests of linkage with Yr28 data are unlikely to be available in the foreseeable future.
Genotype lists: {1325,970}.

Temporary designations:
YrA. Refers to a phenotype specificity that appears to be controlled by complementary genes {1563}. v: Avocet* {*= heterogeneous}; Anza = Karamu = Mexicani =T4 = WW15; Banks*; Condor*; Cocamba; Egret*; Inia 66; Lerma Rojo 64; Lerma Rojo 64A; Nainari 60; Nuri 70; Sanda 73; Sonalika; Zaminder 80. Condor selection P44 Yr6*; Pari 73 Yr6; Saric 70 Yr6; Yecora 70 Yr6 {1563}.

YrCv {939}. 2BS {939}. i: Avocet S*4/Carstens V {970}; Cook*6/Carstens V {970}. v: Caribo {1430}; Cyrano {1430}; Okapi {1430}. Felix Yr3 {1430}. Zdar Yr3a Yr4a {067}.
North American workers {181,186,184} allocated a number of temporary designations to uncatalogued genes detected with North American P. striiformis accessions. Druchamp, Yamhill and Stephens were reported to carry "Yr3a or Yr4a" but these genes could not be distinguished with the cultures that were used. Tests with U.K. isolates showed that Yamhill possessed. Sources of additional genes for seedling (designated "12") and adult resistances ("13", "14", "15") are listed in {1430}.

YrCle {186}. 4B {186}. v: Clement Yr9 {186}.
YrD {185}. 6A {185}. v: Druchamp {185,185}.
YrDru {184,185}. 5B {184},6B {185}. v: Druchamp {184,185}.
YrDru2 {184}. 6A {184}. Druchamp {184}.
YrDa1 {186}. 1A {186}. v: Daws YrDa2 {186}.
YrDa2 {186}. 5D {186}. v: Daws YrDa1 {186}.
YrH46. 6A {184}. Hybrid 46 {184}.
Not the same gene as YrDru2 {184}.
YrHVII {186}. 4A {186}. v: Heines VII Yr2 {186}.
YrMin. 4A {184}. Minister {184}.
YrMor {186}. 4B {186}. v: Moro Yr10 {186}.
YrND. 4A {184}. Nord Desprez {184}.
May be the same as YrMin {184}.
YrS {185}. 3B {185}. v: Stephens {185}.
YrSte {185,184}. 2B {185}, 3B {184}. v: Stephens {185}.
YrSte. 2B {184}. Stephens {184}.
YrSte2. 3B {184}. Stephens {184}.
YrTye {186}. 6D {186}. v: Tyee {186}.
YrTr1 {186}. 6D {186}. v: Tres YrTr2 {186}.
YrTr2 {186}. 3A {186}. v: Tres YrTr1 {186}.
YrYam {185}. 4B {185}. v: Yamhill {185}.
YrYam. 4B {184}. Yamhill {184}.
YrV23. 2B {184}. Vilmorin {184}.
Allelic but not the same as YrSte {184}.

Reaction to Pyrenophora tritici-repentis
Disease: Tan spot, yellow leaf spot.
Tan spot produces two types of genetically determined symptoms, viz. extensive chlorosis and tan necrosis. Pathotypes with the ability to inflict tan spot necrosis (TSN} produce a host gene-specific toxin in culture.

1. Insensitivity to tan spot toxin

tsn1 {346}. Insensitivity (disease resistance) is recessive {346}. 5BL {346}. v: Synthetic W-7976 = Cando/R143/Mexicali ‘S’/3/Ae. squarrosa C122. ma: Xbcd1030-5B - 5.7 cM - tsn1 - 16.5 cM - Xwg583-5B {346}.

Tsn1. Sensitive. v: Kulm {346}.

2. Resistance to chlorosis induction

Tsc1 {344}. Resistant is dominant {344}. 1AS {344}. v: Synthetic W7984 {344}.
tsc1 {344}. v: Opata 85 {344}.
Using a QTL approach, Tsc1 was identified with marker Gli-A1 {344}.

Reaction to Schizaphis graminum Rond. (Toxoptera graminum Rond.)
Insect pest: Greenbug

Gb1 {1514}. [gb1 {222}]. Recessive. v: CI 9058 {222}; Dickinson Selection 28A {222}.

Gb2 {1313,1514}. 1A {554} = T1AL.1R#2S {389} v: Amigo CI 17609 {1313}.

Gb3 {624,1514}. 7D {554}. v: Largo CI 17895 {622}.
Lazar et al. {783} found that resistance in Largo and derivatives was controlled by multiallelic complementary genes. On their conclusions, Gb3 would be one of the loci concerned.
Gb4 {523,1514}. v: CI 17959 {903}.

Gb5 {1514,1515}. 7S {391}, T7S#1L.7S#1S-7AS {389}. su: CI 17882, CI 17884, CI 17885 {1515}. In each of these wheats chromosome 7S substitutes for 7A {391}.

Gb6. 1A = T1AL.1R#2S{1151}. v: GRS1201 {1152}; GRS1202 {1152}; GRS1203 {1152}; GRS1204 {1152}; GRS1205 {1152}; see also Pm17 (Reaction to Erysiphe graminis). su: Tx4386 {1150}. ad: Tx4333 {1150}. al: Insave rye.

Reaction to Tilletia caries (D.C.) Tul., T. foetida (Wallr.) Liro, T. controversa
Disease: Bunt, dwarf smut, stinking smut.

Bt1. [M1 {135}]. 2B {1310}. s: CS*7/White Federation 38 {1304}. v: Albit {129}; Banner Berkeley {129}; Federation 41 {137}; Regal {129}; Sherman {137}; White Federation 38 {1166}; White Odessa {137}. Hussar Bt2 {135}. Hyslop Bt4 {733}; McDermid Bt4 {734}; Tyee Bt4 {022}. Columbia Bt6 {1005}. Martin Bt7 {135}; Odessa Bt7 {137}.

Bt2. [H {129}]. v: Canus {137}; Selection PS60-1-1075 {551}; Selection 1403 {137}. Hussar Bt1 {135}.

Bt3. v: Florence {202,203}; Ridit {152,1000,1395}.

Bt4. [T {136}]. 1B {1005,1274,1285}. v: Bison {1285}; Kaw {1285}; Nebred {1285}; Omaha {1285}; Oveson {1235}. Tres {heterogeneous} {023}; Turkey 1558 {137}; Turkey 2578 {137}. Hyslop Bt1 {733}; McDermid Bt1 {734}; Tyee Bt1 {022}. Oro Bt7 {137}; Turkey 3055 Bt7 {137}.

Note: Since Bt4 and Bt6 are very similar, as well as closely linked, only Turkey 3055 should be used as a definite source of Bt4 and, Rio should be used as the source of Bt6.

Bt5. 1B {1001}. v: Hohenheimer {397}; Selection R60-3432 {551}.

Bt6 [R {1418}]. 1B {1005}. v: Rio {1418}; Turkey 10095 & 10097 {053}. Columbia Bt1 {1005}.

Note: Since Bt4 and Bt6 are very similar, as well as closely linked, only Turkey 3055 should be used as a definite source of Bt4 and, Rio should be used as the source of Bt6.

Bt7. [M2 {1275}]. 2D {1000}. s: CS*7/Cheyenne 2D {1000}. v: Baart {1275}; Cheyenne {1000}; Federation {1275}; Gallipoli {1000}; Onas {1275}; Ranee {1000}; Selection 1833 {556}. Martin Bt1 {137}; Odessa Bt1 {137}. Oro Bt4 {1000}; Turkey 3055 Bt4 {1000}. CI 7090 Bt9 {1000}.

Bt8 {1558}. v: PI 178210 {1558}; Yayla 305 {1558}.

Bt9 {1006}. v: PI 166910 {1006}; PI 166921 {1006}; PI 167822 {1006}; Selection M69-2073 {551}. CI 7090 Bt7 {1000}. Jeff Bt10 {1436}; PI 178383 Bt10 {1006}; Ranger Bt10 {1438}.

Bt10 {1004}. v: Fairview {1183}; PI 116301 {1004}; PI 116306 {1004}; Selection M69-2094 {551}. Jeff Bt9 {1436}; PI 178383 Bt9 {1000}; Ranger Bt9 {1438}. Others {239}. ma: Bt10 was completely linked with a 590 bp fragment produced by UBC primer 196 {239}. RAPD - 1.5 + 1.5cM - Bt10 {763}.

Reaction to Tilletia indica Mitra
Disease: Karnal bunt.

Kb1 {394}. v: Chris {394}. CMH77.308 Kb2 {394}.

Kb2 {394}. v: PF7 113 {394}. CMH77. 308 Kb1 {394}. Shanghai #8 Kb4 {394}.

Kb3 {394}. v: Amsel {394}.

Kb4 {394}. v: Shanghai #8 Kb2 {394}.

Kb5 {394}. Recessive {394}. v: Pigeon Kb6 {394}.

Kb6 {394}. Recessive {394}. v: Pigeon K65 {394}.

Reaction to Ustilagotritici {Pers.} Rostrup
Disease: Loose smut.

Ut1 {1073}. v: Florence/Aurore {1073}; Renfrew {1073}; Red Bobs {1074}.

Ut2 {1073}. v: Kota {1073}; Little Club {1073}.

Ut3 {1074}. v: Carma {1074}.

Ut4 {1074}. v: Thatcher/Regent {1074}.
Temporary designation:

Ut-x {1164}. 2BL{1164}. v: Biggar BSR {1164}.
ma: Xcrc4-2B - 14 cM - Ut-x - 10 cM - Xabc153-2B.2 {1164}. Xcrc4-2B.2 (Syn. Xcrc4.2 ) is a SCAR.

Resistance to colonization by Eriophyes tulipae (Aceria tulipae)

Mite pest: Wheat curl mite.
Eriophyes tulipae is the vector of wheat streak mosaic virus (WSMV) and the wheat spot mosaic agent (WSpM).

Curl mite colonization

Cmc1 {1467}. 6DS {1576}. v: Ae. squarrosa CI4/Novamichurinka (= AC PGR 16635) {1467}.

Cmc2 {1573}. Derived from Agropyron elongatum.

6A = T6AS.6Ae#2S {389}.
5B = T5BL.6Ae#2S {389}. v: 875-94-2 {389}.
6D {1575} = T6DL.6Ae#2S {1575,389}. Tr: Rescue Derivative. su: Cadet 6Ae#2(6A) {1575}; Cadet 6Ae#2(6D) {1574}; Rescue 6Ae#2(6A) {1574}; Rescue 6Ae#2(6B) {1574}; Rescue
6Ae#2(6D). ad: Cadet + mono-6Ae#2 {1574}; Rescue + 6Ae#2 {1574}.

Reaction to Wheat Streak Mosaic Virus
Wsm1 {379,440}. Derived from Ag. intermedium.
4A {800} = T4AL.4Ai#2S {391}. i: Karl*4/CI 17884 = PI 583794 = KS93WGRC27 {440}. v: CI 17766 = B-6-37-1 {391,800,1543}.
4D = T4DL.4Ai#2S} {391,389}. v: CI 17884 {391}; KS90H445 {391}; KS90H450 {391}.
T6AS.4Ai#2L + T6AL-4Ai#2S {389}. v: CI17883 {389}.
4Ai#2 {391}. ad: CI 17881, CI 17886 {391}. su: 4Ai#2(4A) CI 15092 {391}. 4Ai#2(4D) CI 17882 CI 17885 {391}.
ma: Wsm1 cosegregated with a STS amplified by the primer set STSJ15 {1456}.
Wsm1 is located in 4Ai#2S. CI 17882, CI 17884, CI 17885 and KS90H445 also carry a 7S chromosome substituting for 7A (See Reaction to Schizaphis graminum).

Reaction to Xanthomonas campestris pv. undulosa
Disease: Bacterial leaf streak

Bls1 {244}. v: Pavon Bls2 {244}. Mochis T88 Bls3 Bls4 {244}. Angostura F88 Bls5 {244}.

Bls2 {244}. v: Pavon Bls1 {244}.

Bls3 {244}. v: Mochis T88 Bls1Bls4 {244}.

Bls4 {244}. v: Mochis T88 Bls1Bls3 {244}.

Bls5 {244}. v: Turnco F88 {244}. Angostura F88 Bls1 {244}.
bls1 bls2 bls3 bls4 bls5: Alondra {244}.