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Stan D. Wullschleger

Stan D. Wullschleger
Group Leader

Plant Systems Biology Group
Environmental Sciences Division
Oak Ridge National Laboratory
Oak Ridge, TN 37831-6422
Phone:  (865)574-7839
Fax:  (865)576-9939
wullschlegsd@ornl.gov

Research Interests

  • Physiological response of plants to global environmental change
  • Whole-tree water use and environmental control of canopy transpiration
  • Growth and maintenance respiration of leaves and woody tissues
  • Plant water relations
  • Carbon sequestration in plants and soils of terrestrial ecosystems
  • Individual-based modeling of forest succession
  • Physiological genetics
  • Advanced instrumentation for the ecological sciences

Background/Education [ Top ]

Education:

  • Ph.D., Crop Physiology, University of Arkansas, Fayetteville, AR – 1990
  • M.S., Tree Physiology, Colorado State University, Ft. Collins, CO –1982
  • B.S., Forest Management, Colorado State University, Ft. Collins, CO –1979

Experience:

  • Science Leader, Ecosystem and Global Change Science (1999-present), Environmental Sciences Division, Oak Ridge National Laboratory, Oak Ridge, TN.
  • Staff Scientist, (1992-present), Environmental Sciences Division, Oak Ridge National Laboratory, Oak Ridge, TN.
  • Alexander Hollaender Distinguished Postdoctoral Fellow (1990-1992), Environmental Sciences Division, Oak Ridge National Laboratory, Oak Ridge, TN.
  • Research Assistant (1985-1990), Department of Agronomy, University of Arkansas, Fayetteville, AR.
  • Plant Physiologist (1982-1985), USDA-ARS, Crops Research Laboratory, Fort Collins, CO.
  • Research Associate (1981-1982), Department of Botany, Colorado State University, Fort Collins, CO.
  • Graduate Research Assistant (1979-1981), Department of Forest and Wood Sciences, Colorado State University, Fort Collins, CO.

Current Research Projects [ Top ]

Throughfall Displacement Experiment
Michael Huston (PI), Oak Ridge National Laboratory; Stan Wullschleger

This project is part of a larger activity taking place on Walker Branch Watershed, one that in total seeks to examine the biochemical, physiological and ecological response of an eastern temperate forest to changes in precipitation. Our particular sub-project focuses on water use of seedlings, saplings and mature trees, and their respond to alterations in the hydrologic cycle.

Free-Air CO2 Enrichment (FACE) in a Closed-Canopy Forest
Richard J. Norby (PI), Oak Ridge National Laboratory; Nelson Edwards, Carla Gunderson, Paul Hanson, Tony King, Gerry O’Neill, Mac Post, Stan Wullschleger, Dev Joslin and Dale Johnson

The goal of this field experiment is to understand how the eastern deciduous forest will be affected by CO2 enrichment of the atmosphere, and to identify important feedbacks from the forest to the atmosphere. This will be approached by measuring the integrated response of an intact forest ecosystem to elevated CO2 concentration, with a focus on stand-level mechanisms.

Improving the Productivity and Sustainability of the Bioenergy Crop Switchgrass
Stan Wullschleger (PI), Oak Ridge National Laboratory; Chuck Garten and Lee Gunter

This project is part of the larger Biofuels Feedstock Development Program, which is a mission-oriented program of research and analysis whose goal is to develop and demonstrate environmentally acceptable crops and cropping systems for producing large quantities of low-cost, high-quality biomass feedstocks. Our herbaceous bioenergy crop project is aimed at understanding the physiology of switchgrass and important relationships between sustainable yield, soil carbon and nitrogen cycling, and genetic diversity.

Productivity of Intensively Managed Hardwood Plantations
Jerry Tuskan (PI), Oak Ridge National Laboratory; Paul Hanson, Tim Tschaplinski and Stan Wullschleger

The objectives of this study are to quantify the environmental and soil-based constraints (nutrients and water) of productivity in cottonwood, sweetgum and sycamore growing in the southeastern United States. Our focus in this project is on above- and below-ground carbon allocation, physiological assessments of photosynthesis and respiration, whole-tree water use, and biochemical basis for osmotic adjustment and drought tolerance.

Modeling the Global Carbon Cycle
Mac Post (PI), Oak Ridge National Laboratory; Tony King and Stan Wullschleger

The goal of this project is to understand the role of global terrestrial ecosystems carbon exchanges with the atmosphere and oceans in relation to the rising atmospheric CO2 concentration and altered temperature, precipitation, and nitrogen deposition. Chieving this goal requires models that integrate understanding of terrestrial processes involving physiological, whole plant, and ecosystems processes and environmental feedbacks to determine carbon balances of the earth's ecosystems at sufficient temporal and spatial resolution to express mechanistic controls on exchanges of CO2 with the atmosphere.

Carbon Allocation and Partitioning in Woody Plants
Jerry Tuskan (PI), Oak Ridge National Laboratory; Stan Wullschleger, Tim Tschaplinski, Lee Gunter and Mark Davis (NREL)

A joint ORNL/National Renewable Energy Laboratory (NREL) Laboratory Director’s Research and Development (LDRD) project that is using Populus as a model woody perennial to obtain estimates of QTLs for carbon allocation to stems and structural roots. The specific objective of the project is to discover the genetic basis for mechanisms controlling the quantity and quality of photosynthate allocation into secondary cell walls of woody plants as a means of understanding the biological processes that underlie the production and use of biomass for bio-energy and bio-based products.

Publications [ Top ]

Global Environmental Change

Gunderson, C.A., R.J. Norby and S.D. Wullschleger. 2000. Acclimation of photosynthesis and respiration to simulated climatic warming in northern and southern populations of Acer saccharum Marsh.: Laboratory and field evidence. Tree Physiology 20:87-96.

Drake, B.G., J. Azcon-Bieto, J. Berry, J. Bunce, P. Dijkstra, J. Farrar, R. Gifford, M. Gonzalez-Meler, G. Koch, H. Lambers, J. Siedow and S. Wullschleger. 1999. Does elevated atmospheric CO2 concentration inhibit mitochondrial respiration in green plants? Plant Cell Environment 22:649-657.

Norby, R.J., S.D. Wullschleger, C.A. Gunderson, D.W. Johnson and R. Ceulemans. 1999. Tree responses to rising CO2 in field experiments: Implications for the future forest. Plant Cell Environment 22:683-714.

Wullschleger, S.D., P.J. Hanson and T.J. Tschaplinski. 1998. Whole-plant water flux in understory red maple exposed to altered precipitation regimes. Tree Physiology 18:71-79.

Wullschleger, S.D., R.J. Norby, J.C. Love and C.D. Runck. 1997. Energetic cost of tissue construction in yellow-poplar and white oak saplings exposed to long-term CO2 enrichment. Annals of Botany 80:289-297.

Wullschleger, S.D., R.J. Norby and C.A. Gunderson. 1997. Forest trees and their response to atmospheric CO2 Enrichment - A Compilation of Results. pp. 79-100. In: Advances in Carbon Dioxide Effects Research, (eds.) L.H. Allen, Jr., M.B. Kirkham, D.M. Olszyk and C.E. Whitman. American Society of Agronomy Special Publication No. 61, Madison, WI.

Norby, R.J., N.T. Edwards, J.S. Riggs, C.H. Abner, S.D. Wullschleger, C.A. Gunderson, E.G. O'Neill. 1997. Temperature-controlled open-top chambers for global change research. Global Change Biology 3:259-267.

Wullschleger, S.D., P.J. Hanson and G.S. Edwards. 1996. Growth and maintenance respiration in leaves of northern red oak seedlings and mature trees after three years of ozone exposure. Plant Cell Environment 19:577-584.

Kelly, J.M., L.J. Samuelson, G. Edwards, P.J. Hanson, D. Kelting, A. Mays and S.D. Wullschleger. 1995. Are seedlings reasonable surrogates for trees? An analysis of ozone impacts on Quercus rubra. Water, Air and Soil Pollution 85:1317-1324.

Norby, R.J., S.D. Wullschleger, and C.A. Gunderson. 1996. Tree responses to elevated CO2 and the implications for forests. In: Carbon Dioxide and Terrestrial Ecosystems. Eds. G.W. Koch and H.A. Mooney. Pg. 1-21. Physiological Ecology Series. Academic Press.

Norby, R.J., S.D. Wullschleger, C.A. Gunderson amd C.T. Nietch. 1995. Increased growth efficiency in Quercus alba L. trees in a CO2-enriched atmosphere. New Phytologist 131:91-97.

Norby, R.J., E.G. O'Neill and S.D. Wullschleger. 1995. Belowground responses to atmospheric carbon dioxide in forests. pg. 397-418. In: W.W. McFee and J.M. Kelly (eds.) Carbon Forms and Functions in Forest Soils. Soil Science Society of America, Madison, WI.

Wullschleger, S.D., R.J. Norby and P.J. Hanson. 1995. Growth and maintenance respiration in stems of Quercus alba after four years of CO2 enrichment. Physiologia Plantarum 93:47-54.

Wullschleger, S.D., J.P. Lynch and G.M. Berntson. 1994. Modeling the below-ground response of plants and soil biota to edaphic and climatic change - What can we expect to gain? Plant and Soil 165:149-160.

Hanson, P.J., L.J. Samuelson, S.D. Wullschleger, T.A. Tabberer and G.S. Edwards. 1994. Seasonal patterns of light-saturated photosynthesis and leaf conductance for mature and seedling Quercus rubra L. foliage - Differential sensitivity to ozone. Tree Physiology 14:1351-1366.

Wullschleger, S.D., L.H. Ziska and J.A. Bunce. 1994. Respiratory responses of higher plants to atmospheric CO2 enrichment. Physiol. Plant. 90:221-229.

Luxmoore, R.J., S.D. Wullschleger and P.J. Hanson. 1993. Forest responses to CO2 enrichment and climate warming. Water, Soil, Air Pollut. 70: 309-323.

Tschaplinski, T.J., R.J. Norby and S.D. Wullschleger. 1993. Responses of loblolly pine seedlings to elevated CO2 and fluctuating water supply. Tree Physiol. 13:283-296.

Gunderson, C.A., R.J. Norby and S.D. Wullschleger. 1993. Foliar gas exchange of two deciduous hardwoods during three years of growth in elevated CO2 : No loss of photosynthetic enhancement. Plant, Cell and Environ. 16:797-807.

Edwards, G.S., S.D. Wullschleger and J.M. Kelly. 1993. Growth and physiology of northern red oak: Preliminary comparisons of mature and seedling responses to ozone. Environ. Pollut. 83: 215-221.

Wullschleger, S.D. and R.J. Norby. 1992. Respiratory cost of leaf growth and maintenance in white oak saplings exposed to atmospheric CO2 enrichment. Can. J. For. Res. 22:1717-1721.

Wullschleger, S.D., R.J. Norby and C.A. Gunderson. 1992. Growth and maintenance respiration in leaves of Liriodendron tulipifera L. saplings exposed to long-term carbon dioxide enrichment in the field. New Phytol. 121:515-523.

Norby, R.J., C.A. Gunderson, S.D. Wullschleger, E.G. O'Neill and M.K. McCracken. 1992. Productivity and compensatory growth responses of yellow-poplar trees to elevated CO2. Nature 357:322-324.

Wullschleger, S.D., R.J. Norby and D.L. Hendrix. 1992. Carbon exchange rates, chlorophyll concentration, and carbohydrate status of two forest tree species to carbon dioxide enrichment. Tree Physiol. 10:21-31.

Global Carbon Cycle

Post, W.M., A.W. King and S.D. Wullschleger. 1997. Historical variations in terrestrial biospheric carbon storage. Global Biogeochemical Cycles 11:99-109.

King, A.W., W.M. Post and S.D. Wullschleger. 1997. The potential response of terrestrial carbon storage to changes in climate and atmospheric CO2. Climatic Change 35:199-227.

Post, W.M., A.W. King and S.D. Wullschleger. 1996. Soil organic matter models and global estimates of soil organic carbon. Eds. (D.S. Powlson, P. Smith, and J.U. Smith), NATO Advanced Science Institute, Series I, vol. 38:201-222.

King, A.W., W.R. Emanuel, S.D. Wullschleger, and W.M. Post. 1995. In search of the missing carbon sink: A model of terrestrial biospheric response to land-use change and atmospheric CO2. Tellus 47B:501-519.

Wullschleger, S.D., W.M. Post and A.W. King. 1995. On the potential for a CO2 fertilization effect in forests - Estimates of the biotic growth factor based on 58 controlled-exposure studies. pg. 85-107. In: G.M. Woodwell and F.T. Mackenzie (eds.) Biotic Feedbacks in the Global Climate System: Will Warming Feed the Warming ? Oxford Press.

Basic Plant Biosciences

Wullschleger, S.D. 1993. Biochemical limitations to carbon assimilation in C3 plants - A retrospective analysis of the A/Ci curves from 109 species. J. Exp. Bot. 44:907-920.

Wullschleger, S.D., P.J. Hanson and C.A. Gunderson. 1992. Assessing the influence of exogenous ethylene on electron transport and fluorescence quenching in leaves of Glycine max. Environ. Exp. Bot. 32:449-455.

Wullschleger, S.D., P.J. Hanson and R.F. Sage. 1992. PHOTOBIO: Modeling the stomatal and biochemical control of plant gas-exchange. J. Nat. Res. Life Sci. Ed. 21:141-145.

Wullschleger, S.D., D.M. Oosterhuis, R.E. Hurron and P.J. Hanson. 1991. Evidence for light-dependent recycling of respired CO2 by the cotton fruit. Plant Physiol. 97:574-579.

Wullschleger, S.D. and D.M. Oosterhuis. 1990. Photosynthetic and respiratory activity of fruiting forms within the cotton canopy. Plant Physiol. 94:463-469.

Wullschleger, S.D. and D.M. Oosterhuis. 1989. The occurrence of an internal cuticle in cotton (Gossypium hirsutum L.) leaf stomates. Environ. Exp. Bot. 29:229-235.

Canopy Transpiration and Forest Water Use

Wullschleger, S.D., P.J. Hanson and D.E. Todd. 2000. Transpiration from a multi-species deciduous forest as estimated by xylem sap flow techniques. Forest Ecology and Management (in press).

Becker, P., F.C. Meinzer and S.D. Wullschleger. 2000. Hydraulic limitation of tree height: a critique. Functional Ecology 14:4-11.

Wullschleger, S.D. and A.W. King. 2000. Radial variation in sap velocity as a function of stem diameter and height in yellow-poplar trees. Tree Physiology 20:511-518.

Edwards, N.T. and S.D. Wullschleger. 2000. Carbon dioxide efflux rates from stems of mature Quercus alba L. and Acer rubrum L. trees do not appear to affected by sap flow. Journal of Sustainable Forestry (in press).

Grimmond C.S.B., P. Hanson, H.P. Schmid, S. Wullschleger and F. Cropley. 2000. Evapotranspiration rates at the Morgan Monroe State Forest AmeriFlux Site: A comparison of results from eddy covariance turbulent flux measurements and sap flow techniques. 15th Conference on Hydrology, American Meteorological Society Long Beach, CA . January 2000, pp. 158-161.

Wullschleger, S.D., F.C Meinzer and R.A. Vertessy. 1998. A critique of whole-plant water use studies in trees. Tree Physiology 18:499-512.

Crop and Soil Science

Bondada, B.R., Oosterhuis, D.M., Wullschleger, S.D., Kim, K.S. and Harris, W.M. 1994. Anatomical considerations related to photosynthesis in cotton (Gossypium hirsutum L.) leaves, bracts, and the capsule wall. J. Exp. Bot. 45:111-118.

Wullschleger, S.D. and D.M. Oosterhuis. 1992. Canopy leaf area development and age-class dynamics in cotton. Crop Sci. 32:451-456.

Wullschleger, S.D. and D.M. Oosterhuis. 1991. Photosynthesis, transpiration, and water-use efficiency of cotton leaves and fruit. Photosynth. 25:505-515.

Kirkpatrick, T.L., D.M. Oosterhuis and S.D. Wullschleger. 1991. Interaction of root-knot nematodes and water stress in two cotton cultivars. J. Nematology 23:462-467.

Oosterhuis, D.M., R.E. Hampton and S.D. Wullschleger. 1991. Water deficit effects on the cotton leaf cuticle and the efficiency of defoliants. J. Prod. Agric. 4:260-265.

Oosterhuis, D.M., H.D. Scott, R.E. Hampton and S.D.Wullschleger. 1990. Physiological response of two soybean [Glycine max (L.) Merr] cultivars to short-term soil flooding. Environ. Exp. Bot. 30:85-92.

Wullschleger, S.D. and D.M. Oosterhuis. 1990. Photosynthesis of individual field-grown cotton leaves during ontogeny. Photosynth. Research 23:163-170.

Wullschleger, S.D. and D.M. Oosterhuis. 1990. Canopy development and photosynthesis of cotton as influenced by nitrogen nutrition. J. Plant Nutr. 13: 1141-1151.

Hampton, R.E., S.D. Wullschleger and D.M. Oosterhuis. 1990. Impact of Verticillium wilt infection on net photosynthesis, respiration, and photorespiration of field-grown cotton. Physiol. Mol. Plant Path. 37:271-280.

Wullschleger, S.D. and D.M. Oosterhuis. 1990. Photosynthetic carbon production and use by developing cotton leaves and bolls. Crop Sci. 30:1259-1264.

Oosterhuis, D.M., S.D. Wullschleger, R.E. Hampton and R.A. Ball. 1990. Physiological response of rice (Oryza sativa L.) to fenoxaprop-induced injury. Weed Sci. 38:459-462.

Fiscus, E.L., S.D. Wullschleger and H.R. Duke. 1984. Integrated stomatal opening as an indicator of water stress in Zea. Crop Sci. 24:245-249.

Herbaceous Bioenergy Crops

McLaughlin, S.B., J. Bouton, D. Bransby, R. Conger, W. Ocumpaugh, D. Parrish, C. Taliaferro, K. Vogel and S. Wullschleger. 1999. Developing switchgrass as a bioenergy crop. In: J. Janick (ed.) Proceeding of the Fourth National New Crops Symposium, Nov. 9-11, 1998. Phoenix, AZ.

McLaughlin, S., J. Bouton, D. Bransby, R. Conger, W. Ocumpaugh, D. Parrish, C. Taliaferro, K. Vogel and S. Wullschleger. 1997. Evaluating and improving switchgrass as a bioenergy crop. pg. 137-143. In: Making a Business from Biomass (eds. R.P. Overend and E. Chornet). Elsevier Science Inc., New York. pp. 869.

Reed, R.L., M.A. Sanderson, S.B. McLaughlin, S.D. Wullschleger, D.I. Bransby, B. Conger, C. Taliaferro, D.D. Wolfe, W.R. Ocumpaugh, M.A. Hussey, J. Read and C.R. Tischler. 1996. Switchgrass as a sustainable bioenergy crop. Bioresouce Technology 56:83-93.

Gunter, L.E., G.A. Tuskan and S.D. Wullschleger. 1996. Diversity among populations of switchgrass based on RAPD markers. Crop Science 36:1017-1022.

Wullschleger, S.D., M.A. Sanderson, S.B. McLaughlin, D.P. Biradar, and A.L. Rayburn. 1996. Photosynthetic rates and ploidy levels among different populations of switchgrass. Crop Science 36:306-312.

Wullschleger, S.D. 1995. Physiological and morphological indicators of yield in forage grasses - A few case studies and a confession. pg. 68-84. In: G.E. Brink (ed.) Proceedings of the 50th Southern Pasture and Forage Crop Improvement Conference. Knoxville, TN, May 23-25, 1994.

Plant Water Relations

Wullschleger, S.D., P.J. Hanson and D.E. Todd. 1996. Measuring stem water content in four deciduous hardwoods with a time domain reflectometer. Tree Physiology 16:809-815.

Wullschleger, S.D. and D.M. Oosterhuis. 1991. Osmotic adjustment and the growth response of seven vegetable crops following water-deficit stress. HortSci. 26:1210-1212.

Oosterhuis, D.M. and S.D. Wullschleger. 1990. Drought tolerance and irrigation scheduling of vegetable crops. Acta Hort. 278:351-358.

West, C.P., D.M. Oosterhuis and S.D. Wullschleger. 1990. Osmotic adjustment in tissues of tall fescue in response to water deficit. Environ. Exp. Bot. 30:149-156.

Wullschleger, S.D. and D.M. Oosterhuis. 1989. Water use efficiency as a function of leaf age and position within the cotton canopy. Plant Soil 120:79-85.

Oosterhuis, D.M. and S.D. Wullschleger. 1987. Water flow through cotton roots in relation to xylem anatomy. J. Exp. Bot. 38:1866-1874.

Oosterhuis, D.M. and S.D. Wullschleger. 1987. Osmotic adjustment in cotton (Gossypium hirsutum L.) leaves and roots in response to water stress. Plant Physiol. 84:1154-1157

Oosterhuis, D.M., M.L. Parker, S.D. Wullschleger and K.S. Kim. 1988. The citrus leaf cuticle in relation to measurement of leaf water potential using thermocouple psychrometers. Plant Cell and Environ. 11:129-135.

Wullschleger, S.D., M.A. Dixon and D.M. Oosterhuis. 1988. Field measurement of leaf water potential with a temperature-corrected in situ thermocouple psychrometer. Plant Cell and Environ. 11:129-135.

Wullschleger, S.D. and D.M. Oosterhuis. 1987. Electron microscope study of cuticular abrasion on cotton leaves in relation to water potential measurements. J. Exp. Bot. 38:660-667.

Wullschleger, S.D. and D.M. Oosterhuis. 1986. A rapid leaf- disc sampler for psychrometric water potential measurements. Plant Physiol. 81:684-685.

Tyree, M.T., E.L. Fiscus, S.D. Wullschleger and M.A. Dixon. 1986. Detection of xylem cavitation in corn under field conditions. Plant Physiol. 82:597-599.

Oosterhuis, D.M. and S.D. Wullschleger. 1989. Psychrometric water potential analysis in leaf discs. pp. 113-133. In: Modern Methods of Plant Analysis, New Series, Volume 9, Gases in Plant and Microbial Cells. H. F. Linskens and J. F. Jackson (eds.). Springer-Verlag, Berlin.

Soil Carbon Sequestration

Garten, C.T., Jr. and S.D. Wullschleger. 2000. Soil carbon dynamics beneath switchgrass as indicated by stable isotope analysis. Journal of Environment Quality 29(2):645-653.

Garten, C.T. and S.D. Wullschleger. 1999. Soil carbon inventories under a bioenergy crop (Switchgrass): Measurement Limitations. Journal of Environmental Quality 28:1359-1365.

Miscellaneous

McLaughlin, S.B., J.D. Joslin, W. Robarge, A. Stone, R. Wimmer and S.D. Wullschleger. 1997. The impacts of acidic deposition and global change on high elevation southern Appalachian spruce-fir forests. In: The Productivity and Sustainability of Southern Forest Ecosystems in a Changing Environment. R.A. Mickler and S. Fox (eds.), Springer-Verlag, New York, pg. 255-277.

McLaughlin, S.B., J.D. Joslin, A. Stone, R. Wimmer and S.D. Wullschleger. 1996. Effects of acid deposition on calcium nutrition and health of Southern Appalachian Spruce-Fir forests. In: Proc. IUFRO Symp. Air Pollution and Multiple Stresses. Eds. R. Cox, K.Percy, K. Jensen and C. Simpson. p. 207-215. Fredericton, New Brunswick, Canada. September 7-9, 1994.

Hanson, P.J., S.D. Wullschleger, S.A. Bohlman and D.E. Todd. 1993. Seasonal and topographic patterns of forest floor CO2 efflux from an upland oak forest. Tree Physiol. 13:1-15.

Wullschleger, S.D., J.E. Cahoon, J.A. Ferguson and D.M. Oosterhuis. 1991. SURFTEMP: Simulation of soil surface temperature using the energy balance equation. J. Agron. Ed. 20:11-15.

Wullschleger, S.D. and C.P.P. Reid. 1990. Implication of ectomycorrhizal fungi in the cytokinin relations of loblolly pine. New Phytol. 116:681-688.

Kidd, F.A., S.D. Wullschleger, K. Dawley and C.P.P. Reid. 1982. Use of Gentamicin in axenic culturing of ectomycorrhizal plants. Appl. Environ. Microbiol. 44:506-508.

Schaffer, B., F.G. Hawksworth, S.D. Wullschleger and C.P.P. Reid. 1983. Cytokinin-like activity related to host reactions to Dwarf mistletoe (Arceuthobium spp.). Forest Sci. 29:66-70.

Awards and Honors [ Top ]

  • Annual Scientific Achievement Award (1998), Environmental Sciences Division, Oak Ridge National Laboratory, Oak Ridge, TN.
  • Alexander Hollaender Distinguished Postdoctoral Fellowship (1990), Oak Ridge Associated Universities, Oak Ridge, TN.
  • Gerald O. Mott Scholarship for Meritorious Graduate Students in Crop Science, Crop Science Society of America (1989).
  • Aubrey E. Harvey Award, Sigma Xi Research Society, University of Arkansas (1988).
  • Outstanding Agronomy Ph.D. Student, Department of Agronomy, University of Arkansas (1987).
  • Hill Memorial Fellowship, College of Forestry and Natural Resources, Colorado State University (1981).
  • Colorado Graduate Scholarship, Graduate School, Colorado State University (1980).

Professional Activities and Affiliations [ Top ]

Activities
Tree Physiology - Editor

Affiliations
Ecological Society of America, American Society of Agronomy, Crop Science Society of America, Southern Branch of the American Meteorological Society.

Useful Links [ Top ]

September 2005

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