Table Al. Trophic data for Coleoptera collected from the Big Beaver Creek Watershed, NOCA, during 1995 and 1996.

Key to Functional Groups (derived from and similar to Parsons et al. 1991):
? = trophic role uncertain, ~ = associated with trophic resource, but may not feed upon it, Ad = larvae fed or provisioned by adults, D= detritivore, Dg = dung feeder, F1 = flower (incl. pollen, nectar) feeder, Fu = fungivore, H = herbivore, Ms = moss feeder, Nf = non-feeding, O = Omnivore, Pa = parasite, Pr = predator, Rt = root feeder, Sd = seed feeder, Sp = sap / exudate feeder, Sv = Scavenger, X = xylophage.

FAMILY	SPECIES	FUNCTIONAL GROUP		COMMENTS
		Adults	Larvae
Amphizoidae	Amphizoa insolens LeConte	Pr	Pr	Prey upon immature/adult stoneflies. Aquatic.
Anthicidae	Anthicus nanus LeConte	Fl?, 0?	Fu?, 0?
Anthicidae	Eurygenius camplanatus LeConte	Fl	D
Anthicidae	Ischalia vancouverensis Harrington	Nf?	~X	Larvae may feed upon fungi in rotting wood.
Buprestidae	Agrilus politus (Say)	Fl	X	Larval hosts include maples and willows.
Byrrhidae	Byrrhus kirbyi LeConte	Ms	Ms	Feed upon mosses
Byrrhidae	Curimopsis albonotata LeConte	Ms	Ms	Feed upon mosses
Byrrhidae	Cytilus alternatus Say	Ms	Ms	Feed upon mosses
Byrrhidae	Exomella pleuralis (Casey)	Ms	Ms	Feed upon mosses
Byrrhidae	Listemus acuminatus (Mannerheim)	Ms	Ms	Feed upon mosses
Byrrhidae	Morychus aeneolus LeConte	Ms	Ms	Feed upon mosses
Byrrhidae	Morychus oblongus LeConte	Ms	Ms	Feed upon mosses
Cantharidae	Cantharis oregonus LeConte	Fl, Pr	Pr	Adults and larvae prey upon small invertebrates.
Cantharidae	Malthodes alexanderi Fender	Pr?	Pr?
Cantharidae	Malthodes sp.	Pr?	Pr?
Cantharidae	Podabrus conspiratus Fall	Pr	Pr	Adults and larvae prey upon small invertebrates.
Cantharidae	Podabrus piniphilus Dejean	Pr	Pr	Adults and larvae prey upon small invertebrates.
Carabidae	Agonum affine Kirby	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Agonum brevicolle Dejean	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Agonum consimile (Gyllenhal)	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Agonum ferruginosum Dejean	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Agonum piceolum LeConte	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Agonum thoreyi Dejean	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Amara littoralis Mannerheim	O	Pr	Adults readily feed upon seeds.
Carabidae	Amara sanjuanensis Hatch	O	Pr	Adults readily feed upon seeds.
Carabidae	Anchomenus quadratus (LeConte)	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Anisodactylus binotatus Fabricius	O	Pr	EXOTIC, Adults feed upon seeds, plant parts.
Carabidae	Apristus constrictus Casey	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion breve (Motschulsky)	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion concretum Casey	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion convexulum Hayward	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion erasum LeConte	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion fortestriatum Motschulsky	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion hesperum Casey	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion inaequale Say	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion incrematum LeConte	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion iridescens LeConte	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion kuprianovi Mannerheim	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion planatum LeConte	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion planiusculum Mannerheim	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion quadrifoveolatum Mannerheim	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion quadrimaculatum dubitans LeConte	Pr / H /Sd	Pr	Adults known to eat young plants and seeds.
Carabidae	Bembidion quadrulum LeConte	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bembidion semipunctatum Donovan	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Blethisa oregonensis LeConte	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Bradycellus conformis Fall	O	Pr	Adults readily feed upon seeds, vegetation.
Carabidae	Bradycellus lecontei Csiki	O	Pr	Adults readily feed upon seeds, vegetation.
Carabidae	Bradycellus nigrinus Dejean	O	Pr	Adults readily feed upon seeds, vegetation.
Carabidae	Calathus fuscipes (Goeze)	Pr / H /Sd	Pr	EXOTIC, Adults known to feed on plants and seeds.
Carabidae	Chlaenius interruptus Horn	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Cicindela depressula Casey	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Cicindela oregona LeConte	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Diplous atterrimus Dejean	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Elaphrus clairvillei Kirby	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Elaphrus purpurans Hausen	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Harpalus cordifer Notman	O	Pr	Adults extensively herbivorous - seeds, etc.
Carabidae	Harpalus somnulentus Dejean	O	Pr	Adults extensively herbivorous - seeds, etc.
Carabidae	Leistus ferruginosus Mannerheim	Pr	Pr	Adults specialized predators of springtails.
Carabidae	Loricera decempunctata Eschscholtz	Pr	Pr	Adults specialized predators of springtails.
Carabidae	Nebria gebleri cascadensis Kavanaugh	Pr	Pr	Adults & larvae feed upon small invertebrates.
Carabidae	Nebria mannerheimi Fischer	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Nebria sahlbergi Fischer	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Notiophilus sylvaticus Eschscholtz	Pr	Pr	Adults prey upon springtails, mites.
Carabidae	Opisthius richardsoni Kirby	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Patrobus fossifrons dimorphicus Darlington	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Pterostichus adstrictus Eschscholtz	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Pterostichus castaneus (Dejean)	Pr / Sd?	Pr	Adults may feed upon seeds, as do congeners.
Carabidae	Pterostichus herculaneus Mannerheim	Pr / Sd	Pr	Adults known to feed upon seeds.
Carabidae	Pterostichus neobrunneus Lindroth	Pr / Sd?	Pr	Adults may feed upon seeds, as do congeners.
Carabidae	Pterostichus riparius Dejean	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Scaphinotus angulatus Harris	Pr	Pr	Specialized predators of slugs & snails.
Carabidae	Scaphinotus angusticollis Mannerheim	Pr	Pr	Prey on slugs, snails, thought to eat other invertebrates.
Carabidae	Scaphinotus marginatus Fischer	Pr	Pr	Prey on slugs, snails, thought to eat other invertebrates.
Carabidae	Synuchus impunctatus Say	Pr / Sd	Pr	Esp. subcortical. Adults known to eat seeds.
Carabidae	Trechus chalybeus Dejean	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Trechus oregonensis Hatch	Pr	Pr	Adults & larvae prey upon invertebrates.
Carabidae	Trichocellus cognatus Gyllenhal	Pr	Pr	Adults & larvae prey upon invertebrates.
Cerambycidae	Brachyleptura dehiscens (LeConte)	Fl	X	Larval hosts are recently dead conifers.
Cerambycidae	Leptura obliterata Haldeman	Fl	X	Larval hosts are dead and dying conifers.
Cerambycidae	Plectura spinicauda Mannerheim	Fl	X	Larval hosts include hardwoods and conifers.
Cerambycidae	Stenocorus flavolineatus (LeConte)	Fl	X	Larval hosts are probably hardwoods.
Cerambycidae	Xestoleptura crassipes (LeConte)	Fl	X	Larvae occur in well-decayed conifers.
Cerambycidae	Xestoleptura tibialis LeConte	Fl	X	Larvae occur in well-decayed conifers.
Chrysomelidae	Altica corni Woods	H	H	Recorded from Cornus pubescens.
Chrysomelidae	Altica tombacina Mannerheim	H	H	Recorded from dogwood, fireweed.
Chrysomelidae	Chaetocnema irregularis LeConte	H	Rt	Recorded from a wide variety of plants.
Chrysomelidae	Chrysomela mainensis Bechyne	H	H	Breeds & feeds upon alder foliage.
Chrysomelidae	Crepidodera nana (Say)	H	Rt	Hosts are cottonwood, willow, etc.
Chrysomelidae	Hippuriphila mancula LeConte	H	H	Host is Equisetum. Larvae inside stems.
Chrysomelidae	Macrohaltica ambiens (LeConte)	H	H	Adults & larvae feed on alder leaves.
Chrysomelidae	Macrohaltica caurina Blake	H	H	Adults recorded from Ribes.
Chrysomelidae	Plateumaris nitida Germar	H	H	Water lilies, aquatic plants. Larvae aquatic.
Chrysomelidae	Pyrrhalta punctipennis Mannerheim	H	H	Feed upon willow.
Chrysomelidae	Pyrrhalta spiraeophila Hatch & Beller	H	H	Feed upon spirea.
Ciidae	Cis americanus Mannerheim	Fu	Fu	Feed upon polypore fungi.
Ciidae	Cis maritimus Hatch	Fu	Fu	Feed upon fungi.
Ciidae	Octotemnus laevis Casey	Fu	Fu	Known from many polypore fungi
Coccinellidae	Hippodamia washingtoni Timberlake	Pr	Pr	Prey upon aphids
Coccinellidae	Scymnus caurinus Horn	Pr	Pr	Prey upon mealybugs, scales, aphids, & mites.
Coccinellidae	Stethorus punctum picipes Casey	Pr	Pr	Prey upon mites, incl. spider mites.
Colydiidae	Lasconotus vegrandis Horn	Pr	Fu?, Pr	Eat subcortical invertebrates. Young larvae eat fungi.
Corylophidae	Orthoperus scutellaris LeConte	Fu	Fu	Adults & larvae feed upon fungal spores.
Cryptophagidae	Anchicera ephippiata Zimmerman	Fu?	Fu?	Eat fungi & molds in litter, nests, etc.?
Cryptophagidae	Anchicera kamtschatica Motschulsky	Fu?	Fu?	Eat fungi & molds in litter, nests, etc.?
Cryptophagidae	Anchicera ochracea Zimmerman	Fu?	Fu?	Eat fungi & molds in litter, nests, etc.?
Cryptophagidae	Anchicera postpallens Casey	Fu?	Fu?	Eat fungi & molds in litter, nests, etc.?
Cryptophagidae	Antherophagus ochraceus Melsheimer	Fl	D?	Larvae occur in bumblebee nests.
Cryptophagidae	Atomaria constricta Casey	Fu	Fu	Eat fungi & molds in litter, nests, etc.
Cryptophagidae	Atomaria quadricollis Casey	Fu	Fu	Eat fungi & molds in litter, nests, etc.
Cryptophagidae	Caenoscelis ferruginea (Sahlberg)	Fu	Fu	Adults & larvae eat molds and fungal spores.
Cryptophagidae	Cryptophagus cellaris (Scopoli)	Fu	Fu	Adults & larvae eat molds and fungal spores.
Cryptophagidae	Cryptophagus confertus Casey	Fu	Fu	Eat fungi & molds in litter, nests, etc.
Cryptophagidae	Cryptophagus lapponicus Gyllenhal	Fu	Fu	Eat fungi & molds in litter, nests, etc.
Cryptophagidae	Cryptophagus tuberculosus Maklin	Fu	Fu	Eat fungi & molds in litter, nests, etc.
Cryptophagidae	Henotiderus lorna Hatch	Fu?	Fu?	Eat fungi & molds in litter, nests, etc.?
Curculionidae	Baris sparsa LeConte	H	H, Rt	Larvae stem & root feeders upon Asteraceae.
Curculionidae	Cryptorhynchus lapathi Linnaeus	H	X	EXOTIC. On Populus, Salix.
Curculionidae	Geoderces horni Van Dyke	H	Rt	Often a pest of caneberries & strawberries.
Curculionidae	Lepesoma (= Dyslobus) lecontei Casey	H	Rt	Adults eat conifer flowers and young cones.
Curculionidae	Lepesoma (= Dyslobus) verrucifera Casey	H	Rt	Adults feed upon foliage.
Curculionidae	Rhyncolus brunneus Mannerheim	X	X	Under bark of dead conifers.
Curculionidae	Steremnius carinatus Boheman	Rt	Rt	Conifer root feeder, girdler. Vector of black-stain root disease.
Curculionidae	Sthereus (= Lobosoma) horridus (Mannerheim)	D?	D?	Adults in litter, on bark, or in logs.
Dytiscidae	Agabus anthracinus Mannerheim	Pr / Sv	Pr	Prey upon small aquatic invertebrates
Dytiscidae	Agabus austinii Sharp	Pr	Pr	Prey upon small aquatic invertebrates
Dytiscidae	Agabus strigulosus (Crotch)	Pr	Pr	Prey upon small aquatic invertebrates
Dytiscidae	Agabus tristis Aube	Pr / Sv	Pr	Prey upon small aquatic invertebrates
Dytiscidae	Agabus sp. (female)	Pr	Pr	Prey upon small aquatic invertebrates
Dytiscidae	Dytiscus sp. (larva)	Pr	Pr	Prey upon aquatic invertebrates/vertebrates
Dytiscidae	Graphoderus perplexus Sharp	Pr	Pr	Hatch (1953) says "rare"
Dytiscidae	Hydroporus pacificus Fall	Pr	Pr	Prey upon small aquatic invertebrates
Dytiscidae	Hydroporus sp.	Pr	Pr	Prey upon small aquatic invertebrates
Dytiscidae	Rhantus suturellus (Harris)	Pr / Sv	Pr	Prey upon small aquatic invertebrates
Elateridae	Agriotes ferrugineipennis LeConte	Fl, H	Rt	Larvae feed upon grasses.
Elateridae	Ampedus carbonicolor (Eschscholtz)	Fl	Pr / D	Larvae subcortical predators of invertebrates.
Elateridae	Ampedus rhodopus LeConte	Fl, H	Pr, SV	Larvae also eat bacterial & fungal slimes.
Elateridae	Athous rufiventris Eschscholtz	Fl, H	Pr	Adults nocturnal.
Elateridae	Athous vittiger LeConte	Fl, H	Pr	Adults nocturnal.
Elateridae	Cardiophorus amplicollis Motschulsky	Fl, H	Pr	Associated with sandy areas.
Elateridae	Cardiophorus propinquus Hatch	Fl, H	Pr	Associated with sandy areas.
Elateridae	Ctenicera aeripennis (Kirby)	Fl, H	Pr	Larvae are facultative phytophages.
Elateridae	Ctenicera angusticollis Mannerheim	Fl, H	Pr	Larvae are facultative phytophages.
Elateridae	Ctenicera opacula (LeConte)	Fl, H	?	Larvae are facultative phytophages.
Elateridae	Ctenicera propola columbiana (Brown)	Fl	Pr	Larvae are litter/soil predators of invertebrates.
Elateridae	Ctenicera resplendens (Eschscholtz)	Fl	Pr	Larvae are litter/soil predators of invertebrates.
Elateridae	Ctenicera suckleyi (LeConte)	Fl	Pr	Larvae are litter/soil predators of invertebrates.
Elateridae	Ctenicera volitans Eschscholtz	Fl, H	Pr	Larvae are facultative phytophages.
Elateridae	Ctenicera umbripennis (LeConte)	Fl	Pr	Larvae are litter/soil predators of invertebrates.
Elateridae	Dalopius maritimus Brown	Fl, H	Rt
Elateridae	Eanus striatipennis Brown	Fl, H	Pr	Known from montane bogs & wet meadows.
Elateridae	Hemicrepidius pallidipennis Mannerheim	Fl, H	Pr	Larvae may be facultative phytophages.
Elateridae	Hypnoidus bicolor Eschscholtz	Fl, H	Pr	Riparian. Sometimes a minor agricultural pest.
Elateridae	Hypolithus dispersus (Horn)	Fl, H	Pr	Riparian, in gravel/sand bars.
Elateridae	Hypolithus musculus (Eschscholtz)	Fl, H	Pr	Riparian, in gravel/sand bars.
Elateridae	Hypolithus nocturnus Eschscholtz	Fl, H	Pr	Riparian.
Elateridae	Hypolithus squalidus LeConte	Fl, H	Pr	Riparian.
Elateridae	Hypolithus sp.	Fl, H	Pr	Riparian, in gravel/sand bars.
Elateridae	Ligmargus funebris Candeze	Fl, H	Pr	Riparian.
Elateridae	Megapenthes caprella (LeConte)	Fl, H	X
Elateridae	Migiwa sp. (= striatulus LeConte?)	Fl, H	Pr
Elateridae	Negastrius ornatus (LeConte)	Fl, H	Pr	Riparian, in gravel/sand bars.
Elateridae	Zorochrus caurinus Horn	Fl, H	Pr	Riparian, in gravel/sand bars.
Endomychidae	Xenomycetes laversi Hatch	Fu	Fu	Adults & larvae eat polyporus fungi.
Erotylidae	Triplax antica LeConte	Fu	Fu	Recorded from Polyporus
Gyrinidae	Gyrinus picipes Aube	Pr / Sv	Pr	Feed at water surface on aquatic invertebrates.
Histeridae	Hypocaccus bigemmeus LeConte	Pr	Pr	Sandy areas near water. Larvae in sand.
Hydraenidae	Hydraena vandykei vandykei d'Orchymont	H, Sv	Pr?, Sv?	Eat aquatic periphyton, etc.
Hydraenidae	Ochthebius cribricollis LeConte	H, Sv	Pr?, Sv?	Eat aquatic periphyton, etc.
Hydrophilidae	Cercyon adumbratum Mannerheim	D, Dg, Sv	D, Dg, Sv	Adults & larvae eat rotting vegetation, etc.
Hydrophilidae	Crenitis paradigma d'Orchymont	Sv	Pr	Larvae prey upon small aquatic invertebrates.
Hydrophilidae	Cymbiodyta acuminata Fall	Sv?	Pr	Larvae prey upon small aquatic invertebrates.
Hydrophilidae	Helophorus auricollis Eschscholtz	D, H	Pr	Adults known to feed upon dead grass & algae.
Hydrophilidae	Megasternum posticatum (Mannerheim)	D / Sv	D / Sv	Adults & larvae feed on decaying vegetation.
Laemophloeidae	Rhinomalus cygnaei (Mannerheim)	Fu?	Fu?	Adults & larvae may eat subcortical fungi.
Lampyridae	Phausis skelleyi Fender	Pr	Pr	Larvae eat slugs, snails, insect larvae.
Latridiidae	Enicmus cordatus Belon	Fu	Fu	Adults and larvae eat slime mold spores.
Latridiidae	Melanopthalma americana Mannerheim	Fu	Fu	Adults & larvae eat fungal spores, esp. molds.
Latridiidae	Melanopthalma distinguenda Comolli	Fu	Fu	Adults & larvae eat fungal spores, esp. molds.
Latridiidae	Melanopthalma gibbosa Herbst	Fu	Fu	Adults & larvae eat fungal spores, esp. molds.
Latridiidae	Steposthethus liratus LeConte	Fu	Fu	Adults & larvae probably eat fungal spores.
Leiodidae	Agathidium californicum Horn	Fu	Fu	Feed upon basidiomycetes & slime molds.
Leiodidae	Agathidium concinnum Mannerheim	Fu	Fu	Feed upon basidiomycetes & slime molds.
Leiodidae	Agathidium contiguum Fall	Fu	Fu	Feed upon basidiomycetes & slime molds.
Leiodidae	Agathidium sp. (near contiguum Fall)	Fu	Fu	Feed upon basidiomycetes & slime molds.
Leiodidae	Agathidium jasperinum Fall	Fu	Fu	Feed upon basidiomycetes & slime molds.
Leiodidae	Anisotoma confusa Horn	Fu	Fu	Adults & larvae obligate slime mold feeders.
Leiodidae	Anisotoma errans Brown	Fu	Fu	Adults & larvae obligate slime mold feeders.
Leiodidae	Catops basilaris Say	Sv/Dg	Sv/Dg	Eat carrion, dung, decaying fungi, etc.
Leiodidae	Catops egenus Horn	Sv/Dg	Sv/Dg	Eat carrion, dung, decaying fungi, etc.
Leiodidae	Catops simplex Say	Sv/Dg	Sv/Dg	Eat carrion, dung, decaying fungi, etc.
Leiodidae	Catops spp.	Sv/Dg	Sv/Dg	Eat carrion, dung, decaying fungi, etc.
Leiodidae	Catoptrichus frankenhaeuseri (Mann.)	Sv	Sv	At carrion, rotten fungi. Adults fall/winter.
Leiodidae	Colon complicatum Hatch	Fu?	Fu?	Adults & larvae eat subterranean fungi?
Leiodidae	Colon inerme Mannerheim	Fu?	Fu?	Adults & larvae eat subterranean fungi?
Leiodidae	Colon magnicolle Mannerheim	Fu?	Fu?	Adults & larvae eat subterranean fungi?
Leiodidae	Colon nevadense Horn	Fu?	Fu?	Adults & larvae eat subterranean fungi?
Leiodidae	Colon schuhi Hatch	Fu?	Fu?	Adults & larvae eat subterranean fungi?
Leiodidae	Colon serripoides Hatch	Fu?	Fu?	Adults & larvae eat subterranean fungi?
Leiodidae	Colon sp.	Fu?	Fu?	Adults & larvae eat subterranean fungi?
Leiodidae	Hydnobius simulator Brown	Fu	Fu	Adults & larvae feed upon subterranean fungi.
Leiodidae	Leiodes alesi Baronowski	Fu	Fu	Adults & larvae eat subterranean fungi.
Leiodidae	Leiodes cascadensis Baronowski	Fu	Fu	Adults & larvae eat subterranean fungi.
Leiodidae	Leiodes lateritia (Mannerheim)	Fu	Fu	Adults & larvae eat subterranean fungi.
Leiodidae	Leiodes puncticollis (Thomson)	Fu	Fu	Adults & larvae eat subterranean fungi.
Leiodidae	Leptinus occidentamericanus Peck	Sv	Sv	Ectoparasite/inquilines of small mammals.
Leiodidae	Nemadus decipiens (Horn)	D / Sv	D / Sv	Adults & larvae eat decaying organic matter.
Leiodidae	Platycholeus opacellus Fall	D?	D?	Termitophilous with dampwood termites.
Lucanidae	Ceruchus striatus LeConte	Nf	X	Larvae eat sapwood of decaying Douglas fir.
Lycidae	Dictyopterus simplicipes Mannerheim	Fl, Pr	D?	Adults feed upon nectar, probably invertebrates.
Melandryidae	Xylita laevigata Hellenius	Fu?	Fu?	Adults and larvae in decaying conifer logs.
Melyridae	Hypebaeus bicolor (LeConte)	Pr ? ~Fl	Pr? / Sv?	Adults in other genera eat small invertebrates.
Mordellidae	Mordella atrata Melsheimer	Fl	Fu?	Larvae of some congeners in shelf fungi.
Nitidulidae	Epuraea avara Randall	Sv?, Sp	Sv?, Sp	At sap, fermenting fruit, flowers, under bark.
Oedemeridae	Calopus angustus LeConte	Fl	Fu?, X?	Larvae in both live and dead trees.
Oedemeridae	Ditylus gracilis LeConte	Nf?	X	Larvae feed upon moist rotten conifer wood.
Oedemeridae	Ditylus quadricollis LeConte	Nf?	X	Larvae feed upon moist rotten conifer wood.
Oedemeridae	Xanthochroa testacea Horn	Nf?	X	Larvae in decaying wood.
Phalacridae	Phalacrus pencillatus Say	Fl, Fu	Fu	Adults, larvae eat smuts, rusts, mildews, etc.
Phalacridae	Stilbus apicalis Melsh.	Fu	Fu	Eat molds & mildews on dead or dying leaves.
Ptiliidae	Acrotrichis cognata Matthews	Fu	Fu	Adults and larvae feed upon fungal spores.
Ptiliidae	Acrotrichis henrici Matthews	Fu	Fu	Adults and larvae feed upon fungal spores.
Ptiliidae	Acrotrichis vicina Matthews	Fu	Fu	Adults and larvae feed upon fungal spores.
Ptiliidae	Ptenidium pusillum Gyllenhal	Fu	Fu	Adults and larvae feed upon fungal spores.
Ptilodactylidae	Araeopidius monachus LeConte	Nf?	D?	Larvae are aquatic, may be detritivores.
Pyrochroidae	Dendroides ephemeroides Mannerheim	Nf	Fu?	Larvae eat fungi under bark, in decayed wood?
Pyrochroidae	Pedilus jonae Young	Fl	D?, Sv?	Larvae associated with decaying vegetation.
Pythidae	Priognathus monilicornis Randall	X?	X?	Adults, larvae under bark, in decaying conifers.
Scarabaeidae	Aegialia lacustris LeConte	D	D	Adults & larvae eat decaying organic matter.
Scarabaeidae	Aegialia opaca Brown	D	D	Adults & larvae eat decaying organic matter.
Scarabaeidae	Aphodius opacus LeConte	Dg	Dg	Recorded from deer dung.
Scarabaeidae	Onthophagus nuchicornis Linnaeus	Dg	Ad-D	EXOTIC. In domestic stock dung.
Scirtidae	Cyphon brevicollis LeConte	Nf?	D	Adults terrestrial, Larvae aquatic detritivores.
Scirtidae	Cyphon padi Linnaeus	Nf?	D	Adults terrestrial, Larvae aquatic detritivores.
Scirtidae	Cyphon variabilis Thunberg	Nf?	D	Adults terrestrial, Larvae aquatic detritivores.
Scolytidae	Gnathotrichus retusus (LeConte)	Fu	Fu	Recorded from alder, conifers. Ambrosia beetles.
Scydmaenidae	Scydmaenus californicus Motschulsky	Pr?	Pr?	Eat mites, springtails, other small invertebrates?
Scydmaenidae	Scydmaenus fuchsi Brendel	Pr?	Pr?	Eat mites, springtails, other small invertebrates?
Scydmaenidae	Veraphis mirabilis Marsh	Pr?	Pr?	Eat mites, springtails, other small invertebrates?
Silphidae	Nicrophorus defodiens Mannerheim	Sv/Pr/Dg	Ad-Sv	Adults: carrion, feces, fly larvae. Larvae: carrion.
Silphidae	Nicrophorus investigator Zetterstedt	Sv/Pr/Dg	Ad-Sv	Adults: carrion, feces, fly larvae. Larvae: carrion.
Silphidae	Thanatophilus lapponicus (Herbst)	Sv	Sv	On carrion. Cold-adapted.
Sphaeritidae	Sphaerites politus Mannerheim	Dg/D/Fu?	Dg?,D?	Adults: decayed plant material, bear dung, fungi.
Staphylinidae	Acidota crenata Fabricius	Pr?	Pr?	Prey upon small invertebrates?
Staphylinidae	Actium barri Park & Wagner	Pr?	Pr?	Eat mites, springtails, other small invertebrates?
Staphylinidae	Actium retractum Casey	Pr?	Pr?	Eat mites, springtails, other small invertebrates?
Staphylinidae	Aleochara bilineata Gyllenhal	Pr	Pa	probable EXOTIC. Prey upon non-adult flies.
Staphylinidae	Aleochara bimaculata Gravenhorst	Pr?	Pr?	probable EXOTIC. In dung, dead vegetation.
Staphylinidae	Aleocharinae			Detritivores, fungivores, predators, etc.
Staphylinidae	Amphicroum maculatum Horn	Fl	?	Adults probably feed upon pollen.
Staphylinidae	Anthobium clarkae Hatch	Pr?	Pr?	Prey upon small invertebrates?
Staphylinidae	Anthobium reflexicolle Casey	Pr?	Pr?	Ex: mouse nests. Prey on small invertebrates?
Staphylinidae	Anthobium sinuosum Hatch	Pr?	Pr?	Prey upon small invertebrates?
Staphylinidae	Atrecus macrocephalus (Nordmann)	Pr	Pr	Adults & larvae feed on small invertebrates.
Staphylinidae	Atrecus punctiventris (Fall)	Pr	Pr	Adults & larvae feed on small invertebrates.
Staphylinidae	Baeocera humeralis Fall	Fu	Fu	Adults & larvae eat slime molds, basidiomycetes.
Staphylinidae	Batrisodes albionicus (Aube)	Pr	Pr	Eat springtails and other small invertebrates.
Staphylinidae	Bisnius hesperidum Smetana	Pr	Pr	Adults & larvae feed on small invertebrates.
Staphylinidae	Bisnius siegwaldi (Mannerheim)	Pr	Pr	Adults & larvae feed on small invertebrates.
Staphylinidae	Bledius cedarensis Hatch	H	H	Adults and larvae feed upon algae and diatoms.
Staphylinidae	Bledius suturalis LeConte	H	H	Adults and larvae feed upon algae and diatoms.
Staphylinidae	Bolitobius kremeri Maklin	Pr	Pr	Mycophilus predator. Eat maggots?
Staphylinidae	Bryophacis discalis (Hatch)	~Fu	~Fu	Associated with fungi, in litter, under bark, etc.
Staphylinidae	Bryophacis punctatissimus Hatch	~Fu	~Fu	Associated with fungi, in litter, under bark, etc.
Staphylinidae	Bryophacis punctulatus Hatch	~Fu	~Fu	Associated with fungi, in litter, under bark, etc.
Staphylinidae	Cupila excavata Park & Wagner	Pr?	Pr?	Eat mites, springtails, other small invertebrates?
Staphylinidae	Cypha crotchii Horn	?	?	Biology unknown.
Staphylinidae	Dianous nitidulus LeConte	Pr	Pr	Adults & larvae feed on small invertebrates.
Staphylinidae	Elonium NEAR barri (Hatch)	Fl	?	Adults probably feed upon pollen.
Staphylinidae	Elonium rugosa (Hatch)	Fl	?	Adults probably feed upon pollen.
Staphylinidae	Empelus brunnipennis Mannerheim	Fu	Fu	Adults & larvae feed upon fungal spores.
Staphylinidae	Erichsonius cinerascens Gravenhorst	Pr?	Pr?	Adults recorded from sphagnum bogs.
Staphylinidae	Eusphalerum fenyesi Bernh.	Fl	?	Adults probably feed upon pollen.
Staphylinidae	Eusphalerum pothos Mannerheim	Fl	?	Adults probably feed upon pollen.
Staphylinidae	Gabrius cushmani Hatch	Pr	Pr	Adults & larvae prey upon small invertebrates.
Staphylinidae	Gabrius picipennis Maklin	Pr	Pr	but also frequent carrion, dung & detritus.
Staphylinidae	Gabrius seattlensis Hatch	Pr	Pr	As for congeners.
Staphylinidae	Gabrius shulli Hatch	Pr	Pr	As for congeners.
Staphylinidae	Hemiquedius fusculus (LeConte)	Pr	Pr	Adults & larvae prey upon small invertebrates.
Staphylinidae	Ischnosoma fimbriatum Campbell	~Fu	~Fu	Adults & larvae are associated with fungi.
Staphylinidae	Ischnosoma pictum (Horn)	~Fu	~Fu	Adults & larvae are associated with fungi.
Staphylinidae	Ischnosoma splendidus (Gravenhorst)	~Fu	~Fu	Adults & larvae are associated with fungi.
Staphylinidae	Lathrobium punctulatum? LeConte	Pr	Pr	Adults & larvae prey upon small invertebrates.
Staphylinidae	Lathrobium vancouveri Casey	Pr	Pr	Adults & larvae prey upon small invertebrates.
Staphylinidae	Lithocaris capitula Casey	Pr?	Pr?	Adults & larvae in decaying plant matter.
Staphylinidae	Lobrathium sp.	Pr	Pr	Adults & larvae prey upon small invertebrates.
Staphylinidae	Lordithon fungicola Campbell	Pr	Pr	Adults and larvae eat fly larvae. Mycophilus.
Staphylinidae	Lordithon poecilus Mannerheim	Pr	Pr	Adults and larvae eat fly larvae. Mycophilus.
Staphylinidae	Lordithon thoracicus Fabricius	Pr	Pr	Adults and larvae eat fly larvae. Mycophilus.
Staphylinidae	Lucifotychus cognatus LeConte	Pr?	Pr?	Eat mites, springtails, other small invertebrates?
Staphylinidae	Lucifotychus impellus Park & Wagner	Pr?	Pr?	Eat mites, springtails, other small invertebrates?
Staphylinidae	Mathrilaeum pictum (Fauvel)	Fl? / Sv?	?	Adults recorded from flowers and carrion.
Staphylinidae	Mathrilaeum subcostatum Maklin	~Fl, Sv?	?	Adults on flowers and carrion.
Staphylinidae	Megarthrus arcuatus Hatch	Fu	Fu	Adults eat Agaricales, larvae eat fungal spores.
Staphylinidae	Megarthrus pictus Motschulsky	Fu	Fu	Adults eat Agaricales, larvae eat fungal spores.
Staphylinidae	Megarthrus sinuaticollis Boisduval & Lacordaire	Fu	Fu	Adults eat Agaricales, larvae eat fungal spores.
Staphylinidae	Microedus austinianus LeConte	?	?	Riparian along montane sand and gravel bars.
Staphylinidae	Microedus laticollis (Mannerheim)	Pr?	Pr?	Adults and larvae may eat small invertebrates.
Staphylinidae	Micropeplus minor Campbell	Fu	Fu	Adults and larvae eat basidiomycetes, molds.
Staphylinidae	Micropeplus nelsoni Campbell	Fu	Fu	Adults and larvae eat basidiomycetes, molds.
Staphylinidae	Mycetoporus americanus Erichson	~Fu	~Fu	Adults and larvae associated with fungi.
Staphylinidae	Mycetoporus bipunctatus Campbell	~Fu	~Fu	Adults and larvae associated with fungi.
Staphylinidae	Mycetoporus pacificus Campbell	~Fu	~Fu	Adults and larvae associated with fungi.
Staphylinidae	Nitidotachinus tachyporoides (Horn)	Pr	Pr	Adults & larvae feed upon small invertebrates.
Staphylinidae	Olophrum consimile Gyllenhal	?	?	Adults: in litter & debris along streams, ponds.
Staphylinidae	Omalium foraminosum Maklin	Pr	Pr	Congener at carrion, decaying plant material.
Staphylinidae	Ontholestes cingulatus Gravenhorst	Pr	Pr	Associated with dung. Feed upon flies?
Staphylinidae	Oropodes dybasi Grigarick & Schuster	Pr?	Pr?	Eat mites, springtails, other small invertebrates?
Staphylinidae	Oropus striatus (LeConte)	Pr?	Pr?	Eat mites, springtails, other small invertebrates?
Staphylinidae	Orus punctatus Casey	Pr?	Pr?	Adults & larvae prey upon small invertebrates?
Staphylinidae	Oxyporus occipitalis Fauvel	Fu	Fu	Eat fleshy fungi (basidiomycetes: Agaricales).
Staphylinidae	Oxytelus laqueatus Marsham	~Dg, Sv?	~Dg, Sv?	From carrion, dung, detritus. Saprophagous.
Staphylinidae	Pelecomalium testaceum Mannerheim	Fl	?	Adults: esp. at skunk cabbage. Probably pollenivores.
Staphylinidae	Philonthus crotchi Horn	Pr	Pr	Adults & larvae prey on mycophagous flies.
Staphylinidae	Philonthus cruentatus (Gmelin)	Pr	Pr	EXOTIC. In dung & carrion. Prey upon flies.
Staphylinidae	Philonthus duplicatus Bernhauer & Schubert	Pr	Pr	Adults & larvae prey on mycophagous flies.
Staphylinidae	Philonthus furvus Nordmann	Pr	Pr	Recorded from carrion. Prey upon flies.
Staphylinidae	Philonthus spiniformis Hatch	Pr	Pr	Adults & larvae feed upon small invertebrates.
Staphylinidae	Philonthus varians Paykull	Pr	Pr	Adults & larvae prey on mycophagous flies.
Staphylinidae	Phlaeopterus frosti Hatch	Pr / Sv?	Pr / Sv?	Congeneric adults seen eating snowfield insects.
Staphylinidae	Proteinus basalis Maklin	Fu/Sv	Fu/Sv	Under carrion, in moss.
Staphylinidae	Proteinus collars Hatch	Fu/Sv	Fu/Sv	Recorded for alder & pine litter
Staphylinidae	Proteinus limbatus Maklin	Fu/Sv	Fu/Sv	Fungi, leaf litter, compost
Staphylinidae	Pseudopsis sulcata Newman	Fu	Fu	Possible EXOTIC. Eat polyporous fungi.
Staphylinidae	Quedius aenescens Maklin	Pr	Pr	Adults & larvae prey upon invertebrates.
Staphylinidae	Quedius breviceps Casey	Pr	Pr	Adults & larvae prey upon invertebrates.
Staphylinidae	Quedius crescenti Hatch	Pr	Pr	Adults & larvae prey upon invertebrates.
Staphylinidae	Quedius fulvicollis (Stephens)	Pr	Pr	Recorded from swamps, marshes and bogs.
Staphylinidae	Quedius griffinae Hatch	Pr	Pr	Adults & larvae prey upon invertebrates.
Staphylinidae	Quedius horni Hatch	Pr	Pr	Adults & larvae prey upon invertebrates.
Staphylinidae	Quedius nevadensis Casey	Pr	Pr	Adults & larvae prey upon invertebrates.
Staphylinidae	Quedius oculeus Casey	Pr	Pr	Adults & larvae prey upon invertebrates.
Staphylinidae	Reichenbachia albionica Motschulsky	Pr?	Pr?	Eat mites, springtails, other small invertebrates
Staphylinidae	Sonoma hespera Park & Wagner	Pr?	Pr?	Eat mites, springtails, other small invertebrates
Staphylinidae	Staphylinus pleuralis LeConte	Pr	Pr	Adults & larvae prey upon invertebrates.
Staphylinidae	Staphylinus rutilicauda Horn	Pr	Pr	Adults & larvae prey upon invertebrates.
Staphylinidae	Stenus juno Fabricius	Pr	Pr	Adults specialized for capturing springtails.
Staphylinidae	Stenus laccophilus Casey	Pr	Pr	Adults specialized for capturing springtails.
Staphylinidae	Stenus maritimus Motschulsky	Pr	Pr	Adults specialized for capturing springtails.
Staphylinidae	Stenus occidentalis Casey	Pr	Pr	Adults specialized for capturing springtails.
Staphylinidae	Stenus plicipennis (Casey)	Pr	Pr	Adults specialized for capturing springtails.
Staphylinidae	Stenus subgriseus Casey	Pr	Pr	Adults specialized for capturing springtails.
Staphylinidae	Subhaida ingrata (Hatch)	?	?	Biology not recorded.
Staphylinidae	Tachinus basalis Erichson	Pr, ~Fu	Pr, ~Fu	From dung, carrion and decaying fungi.
Staphylinidae	Tachinus crotchii Horn	Pr, ~Fu	Pr, ~Fu	Recorded from carrion, dung & fleshy fungi.
Staphylinidae	Tachinus maculicollis Maklin	Pr, ~Fu	Pr, ~Fu	Recorded from carrion, dung & fleshy fungi.
Staphylinidae	Tachinus nigricornis Mannerheim	Pr, ~Fu	Pr, ~Fu	From dung, carrion and decaying fungi.
Staphylinidae	Tachinus semirufus Horn	Pr, ~Fu	Pr, ~Fu	From dung, carrion and decaying fungi.
Staphylinidae	Tachyporus canadensis Campbell	Pr	Pr	Adults from herbage, ant & animal nests.
Staphylinidae	Tachyporus chrysomelinus Linnaeus	Pr	Pr	Adults from herbage, ant & animal nests.
Staphylinidae	Tachyporus jocosus Say	Pr	Pr	Adults from herbage, ant & animal nests.
Staphylinidae	Tachyporus maculicollis Campbell	Pr	Pr	Adults from herbage, ant & animal nests.
Staphylinidae	Tachyporus mexicanus Sharp	Pr	Pr	Adults from herbage, ant & animal nests.
Staphylinidae	Trichophya pilicornis Gyllenhal	~Fu	~Fu	Possible EXOTIC. Associated with fungi.
Staphylinidae	Unamis fulvipes Fall	?	?	Biology unrecorded.
Tenebrionidae	Helops pernitens LeConte	X?	X?	Adults found in decaying logs and wood.
Tenebrionidae	Scaphidema pictum Horn	D	D	Adults and larvae in sandy riparian areas.
Throscidae	Aulonthroscus validus LeConte	~Fl	Pr?, Fu?>	Live in decaying wood. Reportedly predaceous,.
Throscidae	Pactopus hornii LeConte	~Fl	Pr?, Fu?>	Same as above, but in Europe reportedly eating mycorrhizae.
Trogositidae	Temnochila chlorodia Mannerheim	Pr	Pr	Eat subcortical invertebrates, especially scolytids.
Zopheridae	Phellopsis porcata LeConte	Fu	Fu	Feed upon polypore fungi. Also under bark.

http://www.nps.gov/noca/arthropod2-ta1.htm
Last Updated: 25-Nov-1998