Isla del Coco Fishes

Rhincodontidae — Whale shark
Rhincodon typus Smith, 1828 (photos & video, Garrison)

ORDER LAMNIFORMES

Alopiidae — Thresher sharks
Alopias vulpinus (Bonnaterre, 1788)

ORDER CARCHARHINIFORMES

Carcharhinidae — Requiem sharks
Carcharhinus albimarginatus (Rüppell, 1837)
Carcharhinus falciformis (Müller & Henle, 1839) (photos, Garrison)
Carcharhinus galapagensis (Snodgrass & Heller, 1905)
Carcharhinus limbatus (Müller & Henle, 1839)
Galeocerdo cuvier (Peron & LeSueur in: LeSueur, 1822)
Rhizoprionodon longurio (Jordan & Gilbert, 1882)
Triaenodon obesus (Rüppell, 1837)

Sphyrnidae — Hammerhead sharks
Sphyrna lewini (Griffith & Smith, 1834) (photos & video, Garrison)

ORDER RAJIFORMES

Rhinobatidae — Guitarfishes
Rhinobatos planiceps Garman, 1880

ORDER MYLIOBATIFORMES

Dasyatidae — Whiptail stingrays
Taeniura meyeni Müller & Henle, 1841

Myliobatidae — Eagle rays
Aetobatus narinari (Euphrasen, 1790) (photos & video, Garrison)

Rhinopteridae — Cow-nosed rays
Rhinoptera steindachneri Evermann & Jenkins, 1892

Mobulidae — Mantas, devil rays and mobulas
Manta birostris (Walbaum, 1792) (photos, Garrison)
Mobula munkiana Notarbartolo-di-Sciara, 1987 (sight record, unknown)
Mobula tarapacana (Philippi, 1893) (photos, Garrison)

CLASS ACTINOPTERYGII
ORDER ANGUILLIFORMES

Muraenidae — Morays
Anarchias galapagensis (Seale, 1940)
Echidna nebulosa (Ahl, 1789)
Echidna nocturna (Cope, 1872)
Enchelycore octaviana (Myers & Wade, 1941)
Gymnomuraena zebra (Shaw in: Shaw & Nodder, 1797)
Gymnothorax buroensis (Bleeker, 1857)
Gymnothorax castaneus (Jordan & Gilbert, 1883) (photos)
Gymnothorax dovii (Günther, 1870)
Gymnothorax flavimarginatus (Rüppell, 1830)
Gymnothorax javanicus (Bleeker, 1859) (Masuda et al. 1984; photos, Arroyo & Robertson)
Gymnothorax meleagris (Shaw in: Shaw & Nodder, 1795) (Masuda et al. 1984; photos, Arroyo & Howard)
Gymnothorax panamensis (Steindachner, 1876)
Gymnothorax pictus (Ahl, 1789)
Muraena argus (Steindachner, 1870) (photo, Garrison)
Muraena lentiginosa Jenyns, 1842
Scuticaria tigrina (Lesson, 1828) (photo, Garrison; video, Arroyo)
Uropterygius macrocephalus (Bleeker, 1865)
Uropterygius versutus Bussing, 1991

Congridae — Conger and garden eels
Ariosoma gilberti (Ogilby, 1898)
Heteroconger klausewitzi (Eibl-Eibesfeldt & Köster, 1983)

Ophichthidae — Snake eels
Bascanichthys bascanoides Osburn & Nichols, 1916
Callechelys eristigma McCosker & Rosenblatt, 1972
Gordichthys cf. combibus
Herpetoichthys fossatus (Myers & Wade, 1941)
Ichthyapus selachops (Jordan & Gilbert, 1882)
Myrichthys aspetocheiros McCosker & Rosenblatt, 1993
Myrichthys tigrinus Girard, 1859 (photo, Garrison)
Ophichthus rugifer Jordan & Bollman, 1890
Paraletharchus pacificus (Osburn & Nichols, 1916)
Quassiremus evionthas (Jordan & Bollman, 1890) (photos, Garrison)
Scytalichthys miurus (Jordan & Gilbert, 1882)

ORDER GONORYNCHIFORMES

Chanidae — Milkfishes
Chanos chanos (Forsskål, 1775)

ORDER AULOPIFORMES

Aulopidae — Flagfins
Aulopus bajacali Parin & Kotlyar, 1984

Chlorophthalmidae — Greeneyes
Chlorophthalmus mento Garman, 1899

Synodontidae — Lizardfishes
Synodus lacertinus Gilbert, 1890

ORDER OPHIDIIFORMES

Ophidiidae - Cusk-eels
Otophidium indefatigabile Jordan & Bollman, 1890
Petrotyx hopkinsi Heller & Snodgrass, 1903

Carapidae - Pearlfishes
Carapus dubius (Putnam, 1874)
Echiodon exsilium Rosenblatt, 1961
Encheliophis vermicularis Müller, 1842

Bythitidae - Viviparous brotulas
*Ogilbia new species (Lavenberg, ms)

ORDER GADIFORMES

Merlucciidae —Merlucciid hakes
Merluccius angustimaus  Garman, 1899

ORDER LOPHIIFORMES

Lophiidae — Goosefishes
Lophiodes caulinaris (Garman, 1899) (as L. setigerus)

Antennariidae — Frogfishes
Antennarius avalonis Jordan & Starks, 1907
Antennarius coccineus (Lesson, 1831)
Antennarius commersoni (Latreille, 1804) (photos, Garrison)
Antennarius sanguineus Gill, 1863
Antennarius strigatus (Gill, 1863)

Ogcocephalidae — Batfishes
*Ogcocephalus porrectus Garman, 1899
Zalieutes elater (Jordan & Gilbert, 1882)

ORDER MUGILIFORMES

Mugilidae - Mullets
Agonostomus monticola (Bancroft, 1834)
Chaenomugil proboscideus (Günther, 1861)
Mugil curema Valenciennes in: Cuvier & Valenciennes, 1836 (sight record, Garrison)

ORDER ATHERINIFORMES

Atherinopsidae - New World silversides
Atherinella eriarcha Jordan & Gilbert, 1882
Melanorhinus cyanellus (Meek & Hildebrand, 1923)

ORDER BELONIFORMES

Exocoetidae — Flyingfishes
Exocoetus monocirrhus Richardson, 1846
Exocoetus volitans Linnaeus, 1758
Hirundichthys speculiger (Valenciennes, 1847)
Prognichthys sealei Abe, 1965
Prognichthys tringa Breder, 1928

Hemiramphidae — Halfbeaks
Euleptorhamphus viridis (Van Hasselt, 1823)
Hyporhamphus rosae (Jordan & Gilbert, 1880)
Oxyporhamphus m. micropterus (Valenciennes in: Cuvier & Valenciennes, 1847)

Belonidae — Needlefishes
Platybelone argalus pterura (Osburn & Nichols, 1916)
Strongylura exilis (Girard, 1854)
Tylosurus acus melanotus (Bleeker, 1851)
Tylosurus crocodilus fodiator Jordan & Gilbert, 1882

ORDER BERYCIFORMES

Holocentridae — Soldierfishes and squirrelfishes
Myripristis berndti Jordan & Evermann, 1903
Myripristis leiognathus Valenciennes, 1846
Plectrypops lima (Valenciennes in: Cuvier & Valenciennes, 1831)
Sargocentron suborbitalis (Gill, 1863)

ORDER GASTEROSTEIFORMES

Aulostomidae - Trumpetfishes
Aulostomus chinensis (Linnaeus, 1766)

Fistulariidae - Cornetfishes
Fistularia commersonii Rüppell, 1838

Syngnathidae - Pipefishes and seahorses
Bryx veleronis Herald, 1940
Doryrhamphus excisus excisus Kaup, 1856
Hippocampus ingens Girard, 1858

ORDER SCORPAENIFORMES

Scorpaenidae — Scorpionfishes
Pontinus clemensi Fitch, 1955
Pontinus strigatus Heller & Snodgrass, 1903
Scorpaena afuerae Hildebrand, 1946
*Scorpaena cocosensis Motomura, 2004
Scorpaena histrio Jenyns, 1840
Scorpaena mystes Jordan & Starks in: Jordan, 1895
Scorpaena russula Jordan & Bollman, 1890
Scorpaenodes xyris (Jordan & Gilbert, 1882)

Triglidae — Searobins
Bellator loxias Jordan, in: Gilbert, 1897
*Prionotus new species Lavenberg, ms

Peristediidae — Armored searobins
*Peristedion new species Lavenberg, ms

ORDER PERCIFORMES

Serranidae — Groupers and sea basses
Alphestes immaculatus Breder, 1936 (sight record and photos, Garrison)
Cephalopholis panamensis (Steindachner, 1877)
Dermatolepis dermatolepis (Boulenger, 1895)
Epinephelus cifuentesi Lavenberg & Grove in: Heemstra & Randall, 1993
Epinephelus itajara (Lichtenstein, 1822) (photo, Garrison)
Epinephelus labriformis (Jenyns, 1840)
Epinephelus niphobles Gilbert & Starks in: Gilbert, 1897
Liopropoma fasciatum Bussing, 1980
Mycteroperca olfax (Jenyns, 1840)
Paranthias colonus (Valenciennes, 1846)
Pronotogrammus eos Gilbert, 1890
Pronotogrammus multifasciatus Gill, 1863
Pseudogramma thaumasium (Gilbert in: Jordan & Evermann, 1900)
Rypticus bicolor Valenciennes, 1846
Serranus aequidens Gilbert, 1890
*Serranus tico Meisler & Lavenberg, in Allen & Robertson, 1998

Kuhliidae — Flagtails
Kuhlia mugil (Forster in: Bloch & Schneider, 1801)

Priacanthidae — Bigeyes
Cookeolus japonicus (Cuvier in: Cuvier & Valenciennes, 1829) (photo, Garrison)
Heteropriacanthus cruentatus (Lacépède, 1801)
Pristigenys serrula (Gilbert, 1891) (video, Garrison)

Apogonidae — Cardinalfishes
Apogon atradorsatus Heller & Snodgrass, 1903
Apogon dovii Günther, 1861

Malacanthidae — Tilefishes
Caulolatilus affinis Gill, 1865
Caulolatilus hubbsi Dooley, 1978
Malacanthus brevirostris Guichenot, 1848

Echeneidae — Suckerfishes, remoras
Remorina albescens (Temminck & Schegel, 1850)
Remora brachyptera (Lowe, 1839)
Remora osteochir (Cuvier, 1829)
Remora remora (Linnaeus, 1758)

Carangidae — Jacks and pompanos
Alectis ciliaris (Bloch, 1787)
Caranx caballus Günther, 1868
Caranx caninus Günther, 1867 (video, Garrison)
Caranx lugubris Poey, 1860 (photos, Garrison)
Caranx melampygus Cuvier in: Cuvier & Valenciennes, 1833
Caranx orthogrammus (Jordan & Gilbert, 1882) (video, Garrison)
Caranx sexfasciatus Quoy & Gaimard, 1825 (photos, Garrison)
Decapterus macarellus (Cuvier in: Cuvier & Valenciennes, 1833) (photo, Garrison)
Elagatis bipinnulata (Quoy & Gaimard, 1825)
Gnathanodon speciosus (Forsskål, 1775) (sight records, Garrison)
Naucrates ductor (Linnaeus, 1758)
Selar crumenophthalmus (Bloch, 1793)
Seriola rivoliana Valenciennes in: Cuvier & Valenciennes, 1833 (photo, Garrison)
Trachinotus rhodopus Gill, 1863 (video, Garrison)
Trachinotus stilbe (Jordan & McGregor, 1899)
Uraspis helvola (Forster, 1801) (photos & video, Garrison)

Coryphaenidae — Dolphinfishes
Coryphaena equiselis Linnaeus, 1758
Coryphaena hippurus Linnaeus, 1758

Lutjanidae — Snappers
Aphareus furca (Lacépède, 1801)
Hoplopagrus guntheri Gill, 1862
Lutjanus aratus (Günther, 1864) (photos)
Lutjanus argentiventris (Peters, 1869)
Lutjanus guttatus (Steindachner, 1869) (photo, sight record, Garrison)
Lutjanus inermis (Peters, 1869)
Lutjanus jordani (Gilbert in: Jordan & Evermann, 1898)
Lutjanus novemfasciatus Gill, 1862 (photos & video, Garrison)
Lutjanus viridis (Valenciennes, 1846)

Gerreidae — Mojarras
Eucinostomus currani Zahuranec in: Yáñez-Arancibia, 1980

Lobotidae — Tripletails
Lobotes pacificus Gilbert, 1898

Haemulidae — Grunts
Anisotremus caesius (Jordan & Gilbert, 1882)
Anisotremus interruptus (Gill, 1862)
Anisotremus taeniatus Gill, 1861 (sight record, Garrison)

Sciaenidae — Drums
Umbrina xanti Gill, 1862

Mullidae — Goatfishes
Mulloidichthys dentatus (Gill, 1862)

Kyphosidae — Sea chubs
Kyphosus elegans (Peters, 1869)
Sectator ocyurus (Jordan & Gilbert, 1882)

Chaetodontidae — Butterflyfishes
Chaetodon humeralis Günther, 1860
Chaetodon lunula (Lacépède, 1803) (photos)
Johnrandallia nigrirostris (Gill, 1862)
Prognathodes falcifer (Hubbs & Rechnitzer, 1958)

Pomacanthidae — Angelfishes
Holacanthus passer Valenciennes, 1846
Pomacanthus zonipectus (Gill, 1862)

Pomacentridae — Damselfishes
Abudefduf concolor (Gill, 1862)
Abudefduf troschelii (Gill, 1862)
Chromis alta Greenfield & Woods, 1980 (sight record, Lavenberg)
Chromis atrilobata Gill, 1862
Microspathodon bairdii (Gill, 1862)
Microspathodon dorsalis (Gill, 1862)
Stegastes acapulcoensis (Fowler, 1944)
Stegastes arcifrons (Heller & Snodgrass, 1903)
Stegastes flavilatus (Gill, 1862)
Stegastes beebei (Nichols, 1924)

Cirrhitidae — Hawkfishes
Cirrhitichthys oxycephalus (Bleeker, 1855)
Cirrhitus rivulatus (Valenciennes, 1846)
Oxycirrhites typus Bleeker, 1857

Sphyraenidae — Barracudas
Sphyraena idiastes Heller & Snodgrass, 1903 (photos, Garrison)

Polynemidae — Threadfins
Polydactylus approximans (Lay & Bennett, 1839)

Labridae — Wrasses
Bodianus diplotaenia (Gill, 1862)
Decodon melasma Gomon, 1974
Halichoeres adustus (Gilbert, 1890)
*Halichoeres discolor Bussing, 1983
Halichoeres dispilus (Günther, 1864)
Halichoeres nicholsi (Jordan & Gilbert, 1882)
*Halichoeres salmofasciatus Allen & Robertson, 2002
Halichoeres cf. raisneri (collected, Bussing)
Iniistius pavo (Valenciennes in: Cuvier & Valenciennes, 1840) (sight record, Lavenberg, Bussing; photo, Garrison)
Novaculichthys taeniourus (Lacépède, 1801)
Polylepion cruentum Gomon, 1977
Stethojulis bandanensis (Bleeker, 1851)
Thalassoma grammaticum Gilbert, 1890
Thalassoma lucasanum (Gill, 1862)
Thalassoma purpureum (Forsskål, 1775)
Xyrichtys victori Wellington, 1992

Scaridae — Parrotfishes
Scarus compressus (Osburn & Nichols, 1916) (photo, Garrison)
Scarus ghobban Forsskål, 1775
Scarus perrico Jordan & Gilbert, 1882
Scarus rubroviolaceus Bleeker, 1847

Opistognathidae — Jawfishes
Opistognathus panamaensis Allen & Robertson, 1991

Tripterygiidae — Triplefins
*Axoclinus cocoensis Bussing, 1991

Chaenopsidae — Tube blennies
*Acanthemblemaria atrata Hastings & Robertson, 1999
*Chaenopsis new species (Lavenberg, ms)
Coralliozetus boehlkei Stephens, 1963
Coralliozetus springeri Stephens & Johnson in: Stephens, Hobson & Johnson, 1966
Emblemaria nivipes Jordan & Gilbert, 1883

Dactyloscopidae — Sand stargazers
Dactyloscopus pectoralis fallax Dawson, 1975
*Gillellus chathamensis Dawson, 1977

Labrisomidae — Labrisomid blennies
Paraclinus mexicanus (Gilbert, 1904)
Starksia sp.

Blenniidae — Combtooth blennies
Entomacrodus chiostictus (Jordan & Gilbert, 1882)
Hypsoblennius brevipinnis (Günther, 1861)
Ophioblennius steindachneri Jordan & Evermann, 1898
Plagiotremus azaleus (Jordan & Bollman, 1890)

Gobiesocidae - Clingfishes
Arcos rhodospilus (Günther, 1864)
Gobiesox adustus Jordan & Gilbert, 1882
*Gobiesox fulvus Meek, 1907
*Gobiesox woodsi (Schultz, 1944)
*Tomicodon vermiculatus Briggs, 1955

Callionymidae — Dragonets
Synchiropus atrilabiatus (Garman, 1899)

Eleotridae — Sleepers
Eleotris picta Kner, 1863
*Eleotris tubularis Heller & Snodgrass, 1903

Gobiidae — Gobies
Bathygobius ramosus longipinnis Ginsburg, 1947
*Chriolepis atrimelum, Bussing, 1997
*Chriolepis dialepta Bussing, 1990
Coryphopterus urospilus Ginsburg, 1938 (sight record, C. Swift)
*Elacatinus nesiotes Bussing, 1990
Gobulus hancocki Ginsburg, 1938
*Lythrypnus alphigena Bussing, 1990
*Lythrypnus cobalus Bussing, 1990
*Lythrypnus lavenbergi Bussing, 1990
Lythrypnus rhizophora (Heller & Snodgrass, 1903)
*Sicydium cocoensis (Heller & Snodgrass, 1903)

Microdesmidae — Wormfishes
Clarkichthys bilineatus (Clarke, 1936)

Acanthuridae — Surgeonfishes
Acanthurus nigricans (Linnaeus, 1758)
Acanthurus triostegus (Linnaeus, 1758)
Acanthurus xanthopterus Valenciennes in: Cuvier & Valenciennes, 1835
Ctenochaetus marginatus (Valenciennes in: Cuvier & Valenciennes, 1835)
Prionurus laticlavius (Valenciennes, 1846)

Zanclidae — Moorish idol
Zanclus cornutus (Linnaeus, 1758)

Scombridae — Mackerels and tunas
Acanthocybium solandri (Cuvier in: Cuvier & Valenciennes, 1832)
Auxis rochei eudorax Collette & Aadland, 1996
Auxis thazard brachydorax Collette & Aadland, 1996
Euthynnus lineatus Kishinouye, 1920
Katsuwonus pelamis (Linnaeus, 1758)
Sarda orientalis (Temminck & Schlegel, 1844) (photo & video, Garrison)
Scomberomorus sierra Jordan & Starks in: Jordan, 1895
Thunnus albacares (Bonnaterre, 1788 )

Istiophoridae — Billfishes
Istiophorus platypterus (Shaw in: Shaw & Nodder, 1792) (photo, Garrison)
Makaira mazara (Jordan & Snyder, 1901)
Tetrapturus audax (Philippi, 1887)

ORDER PLEURONECTIFORMES

Bothidae — Lefteye flounders
Bothus leopardinus (Günther, 1862)
Bothus mancus (Broussonet, 1782)

Paralichthyidae — Sand flounders
Hippoglossina bollmani Gilbert, 1890
Syacium latifrons (Jordan & Gilbert, 1882)
*Syacium maculiferum (Garman, 1899)

Soleidae — Soles
Aseraggodes herrei Seale, 1940

Cynoglossidae — Tonguefishes
Symphurus varius Garman, 1899

ORDER TETRAODONTIFORMES

Balistidae — Triggerfishes
Balistes polylepis Steindachner, 1876
Canthidermis maculatus (Bloch, 1786)
Melichthys niger (Bloch, 1786)
Melichthys vidua (Richardson, 1845)
Pseudobalistes naufragium (Jordan & Starks in: Jordan 1895) (sight record, Garrison)
Sufflamen verres (Gilbert & Starks, 1904)
Xanthichthys caeruleolineatus Randall, Matsuura & Zama, 1978 (photos & video)
Xanthichthys mento (Jordan & Gilbert, 1882)

Monacanthidae — Filefishes
Aluterus monoceros (Linnaeus, 1758) (photos, Garrison)
Aluterus scriptus (Osbeck, 1765)
Cantherhines dumerilii (Hollard, 1854)

Ostraciidae — Boxfishes
Ostracion meleagris Shaw in: Shaw & Nodder, 1796

Tetraodontidae — Puffers
Arothron hispidus (Linnaeus, 1758)
Arothron meleagris (Lacépède, 1798)
Canthigaster punctatissima (Günther, 1870)
Sphoeroides angusticeps (Jenyns, 1842) (video, Arroyo)
Sphoeroides lobatus (Steindachner, 1870)

Diodontidae — Porcupinefishes
Chilomycterus reticulatus (Linnaeus, 1758) (photos & video, Garrison; photos, Arroyo & Robertson)
Diodon holocanthus Linnaeus, 1758
Diodon hystrix Linnaeus, 1758