Stock number: BGS 351
Locus name: Glossy sheath 1
Locus symbol: gsh1

Previous nomenclature and gene symbolization:

Glaucous sheath = gs (3).
Glossy sheath and spike = gs (23).
Glossy sheath 1 = gs1 (15).
Eceriferum-q = cer-q (5).

Monofactorial recessive (3, 5), except Cer-q.1440 (12).
Located in chromosome 2HS (19, 20), in a subterminal position (18) over 36.5 cM distal from the Eam1 (early maturity 1) locus (1).

A bright, green glossy color is apparent on the spike, leaf sheath, and stem (wax code - - ++) (5, 15). The gsh1 locus is the cer-q component of the complex locus named cer-cqu (2, 14), which controls formation of -diketone lipids in epicuticular wax layer (21). The cer-q mutants affect the -ketoacyl condensing enzyme (16, 22).

A spontaneous mutant in PI 95285 (3), an iodine vapor induced mutant in Bonus (PI 189763) (5).

gsh1.a in PI 95285 (3, 15, 17); gsh1.b in Piroline (PI 539132) (15); gsh1.c in Velvon (CIho 6109) (15, 23); gsh1.y in Minn 549 (15, 17); cer-q.35 (GSHO 438) in Bonus (2, 5, 6); cer-q.42, -q.50, -q.56, -q.103 in Bonus (5, 6); cer-q.82, -q.83, -q.84, -cqu.124, -q.128, -q.131, -q.132, -q.133, -q.141, -q.151 in Bonus, -q.213, -q.217, -q.218, -q.239, -q.241, -q.245, -q.246, -q.261, -q.262, -q.292, -q.295, -q.297, -q.299, -q.310, -q.320, -q.326, -q.327, -q.334, -q.335, -q.341, -q.365, -q.395, -q.396, -q.400, -q.403, -q.411, -cqu.416, -cqu.420, -q.425, -q.440, -q.488, -q.509, -qu.510 in Foma (CIho 11333) (6, 14); cer-q.129, -q.176 in Bonus, -q.483, -q.516, -q.519, -q.524, -q.527, -q.533, -q.536, -q.548, -q.555, -q.557, -q.566, -q.574, -q.579, -q.597 in Foma, -q.675, -q.683, -q.688, -cqu.724, -cqu.733 in Bonus, -q.1016, -q.1018, -q.1040, -q.1061, -q.1063, -q.1064, -q.1065, -q.1068, -q.1070, -q.1073, -q.1074, -q.1083 in Carlsberg II (CIho 10114) (6); cer-q.153, -q.754, -q.755, -q.756, -q.757, -q.759, -q.772, -qu.813, -q.822, -q.827, -q.837, -q.848, -q.876 in Bonus, -q.401 in Foma, -q.1079 in Carlsberg II, -q.1105, -q.1128, -q.1131, -q.1141, -q.1143 in Kristina (7); cer-q.629, -q.634, -q.636, -cq.645, -q.900, -cqu.925, -cqu.944 in Bonus, -q.1158, -q.1167 in Kristina (8); cer-q.1011 in Carlsberg II, -q.1121, -q.1193, -q.1209, -q.1225, -q.1234, -q.1238, -q.1242, -q.1243 in Kristina (9); cer-q.598 in Foma, -q.1260, -q.1281, -q.1283, -q.1285 in Kristina, -q.1320, -q.1330 in Bonus (10); cer-q.1230 in Kristina, -q.1345, -q.1358, -q.1362, -q.1363, -q.1368, -q.1369, -q.1375, -q.1400, -q.1406, -q.1412, -q.1430, -q.1459, -q.1464 in Bonus, -q.1504, -q.1512, -q.1514 in Nordal (11); Cer-q.1440, cer-q.1490, -q.1499, -q.1706, -q.1708, -q.1742, -q.1748, -q.1753 in Bonus, -q.1517, -q.1521, -q.1522, -q.1527, -q.1528, -q.1533, -q.1534, -q.1537, -q.1538, -q.1539, -q.1542, -q.1551, -q.1556, -q.1572 in Nordal (12); cer-q.571 in Foma (13); gsh1.aa (OUM018), gsh1.ad (OUM021), gsh1.ak (OUM028) in Akashinriki (PI 467400, OUJ659), gsh1.an (T119, OUL045) in a tester line, gsh1.ap (PI 95285, OUE093), gsh1.ar (PI 151791, OUE121), gsh1.as (PI 151815, OUE128), gsh1.at (PI 194916, OUE168), gsh1.au (CIho 14093, OUE330), gsh1.av (CIho 14139, OUE370), gsh1.ax (CIho 14094, OUE630), gsh1.ay (CIho 14140, OUE670), gsh1.az (CIho 4361, OUE676) in Ethiopian introductions, gsh1.ba in Bozu Omugi 1 (OUJ655) (4).
Mutant used for description and seed stocks:
gsh1.a in PI 95285 (GSHO 735); gsh1.a in Bowman (PI 483237)*7 (GSHO 1868).

1. Doney, D.L. 1961. An inheritance and linkage study of barley with special emphasis on purple pigmentation on the auricle. M.S. Thesis. Utah State Univ., Logan.
2. Eslick, R.F. 1974. Allele tests of gs and cer mutants. BGN 4:9-11.
3. Immer, F.R., and M.T. Henderson. 1943. Linkage studies in barley. Genetics 28:419-440.
4. Konishi, T. 1973. Genetic analyses of EMS-induced mutants in barley. BGN 3:28-31.
5. Lundqvist, U., and D. von Wettstein. 1962. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. Hereditas 48:342-362.
6. Lundqvist, U., and D. von Wettstein. 1971. Stock list for the eceriferum mutants. BGN 1:97-102.
7. Lundqvist, U., and D. von Wettstein. 1973. Stock list for the eceriferum mutants II. BGN 3:110-112.
8. Lundqvist, U., and D. von Wettstein. 1975. Stock list for the eceriferum mutants III. BGN 5:88-91.
9. Lundqvist, U., and D. von Wettstein. 1977. Stock list for the eceriferum mutants IV. BGN 7:92-96.
10. Lundqvist, U., and D. von Wettstein. 1979. Stock list for the eceriferum mutants V. BGN 9:135-137.
11. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.
12. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.
13. Lundqvist, U., and D. von Wettstein. 1988. Stock list for the eceriferum mutants VIII. BGN 18:88-91
14. Lundqvist, U., P. von Wettstein-Knowles, and D. von Wettstein. 1968. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. II. Hereditas 59:473-504.
15. McProud, W.L., and R.F. Eslick. 1971. Allelism within a series of glossy sheath mutants. BGN 1:31-34.
16. Mikkelsen, J.D. 1984. Biosynthesis of esterified alkan-2-ols and -diketones in barley spike epicuticular wax: synthesis of radioactive intermediates. Carlsberg Res. Commun. 49:391-416.
17. Rasmusson, D.C, and J.W. Lambert. 1965. Inheritance of the glossy-sheath character in barley, Hordeum vulgare L. Crop Sci. 5:251-253.
18. Schondelmaier, J., G. Fischbeck, and A. Jahoor. 1993. Linkage studies between morphological and RFLP markers in the barley genome. BGN 22:57-62.
19. Takahashi, R., J. Yamamoto, S. Yasuda, and Y. Itano. 1953. Inheritance and linkage studies in barley. Ber. Ohara Inst. landw. Forsch. 10:29-52.
20. Tsuchiya, T. 1972. Cytogenetics of telotrisomics in barley. BGN 2:93-98.
21. Wettstein-Knowles, P. von. 1972. Genetic control of -diketone and hydro- -diketone synthesis in epicuticular waxes of barley. Planta 106:113-130.
22. Wettstein-Knowles, P. von. 1992. Molecular genetics of the lipid synthesis in barley. p.753-771. In L. Munck, K. Kirkegaard, and B. Jensen (eds.) Barley Genetics, Vol. II. Proc. Sixth Int. Barley Genet. Symp., Helsingborg, 1991. Sweden. Munksgaard International Publishers Ltd., Copenhagen.
23. Woodward, R.W. 1957. Linkage studies in barley. Agron. J. 49:28-32.

W.L. McProud and R.F. Eslick. 1971. BGN 1:168-169.

J.D. Franckowiak and U. Lundqvist. 1997. BGN 26:292-293.

Stock number: BGS 352
Locus name: Glossy sheath 2
Locus symbol: gsh2

Previous nomenclature and gene symbolization:

Glossy sheath 2 = gs2 (17).
Eceriferum-b = cer-b (7).

Monofactorial recessive (7, 16).
Located in chromosome 3HL (1, 3, 18), about 5.1 cM proximal from the als (absent lower laterals) locus (4, 5) and over 32.5 cM distal from the uzu (uzu) locus (1, 4, 5).

Surface wax coating appears absent on the spike, leaf sheath, and stem (wax code - - ++) (7, 16).

Spontaneous occurrence in Vantage (CIho 7324) (15); an X-ray induced mutant in Maja (PI 184884) (7).

gsh2.d in Vantage, gsh2.e in Ymer (CIho 7275), gsh2.f, gsh2.g in Atlas (CIho 4118), gsh2.h in Klargrin (15); cer-b.2 in Maja (6, 7, 8, 19); cer-b.4, -b.64, -b.66, -b.109 in Bonus (PI 189763) (7, 8); cer-b.79, -b.94, -b.96 in Bonus, -b.209, -b.222, -b.319, -b.336, -b.337, -b.377, -b.418, -b.424, -b.427 in Foma (CIho 11333) (8, 14); cer-b.169, -b.193 in Bonus, -b.412, -b.531, -b.549 in Foma, -b.1037, -b.1059 in Carlsberg II (CIho 10114) (8); cer-b.751, -b.769, -b.791, -b.799, -b.800 in Bonus, -b.1129 in Kristina (9); cer-b.910 in Bonus (10); cer-b.932 in Bonus (11); cer-b.147, -b.182, -b.623 in Bonus, -b.1099, -b.1267, -b.1269 in Kristina (12); cer-b.1699 in Bonus (13); gsh2.ab (OUM019), gsh2.ah (OUM025) in Akashinriki (OUJ659, PI 467400) (3, 5); gsh2.bb in the cross NDB112/Fr926-77 (2).
Mutant used for description and seed stocks:
gsh2.f in Atlas (GSHO 736); cer-b.2 in Maja (GSHO 434; GSHO 1081); gsh2.f in Bowman (PI 483237)*7 (GSHO 1988); cer-b.2 in Bowman*6 (GSHO 1989).

1. Eslick, R.F., and W.L. McProud. 1974. Positioning of genetic male sterile 5 (msg5) on chromosome 3. BGN 4:16-23.
2. Franckowiak, J.D. 1995. (unpublished).
3. Konishi, T. 1970. Genetic analyses of two kinds of EMS-induced mutants of barley. Barley Newsl. 13:57-58.
4. Konishi, T. 1973. Genetic analyses of EMS-induced mutants in barley. BGN 3:28-31.
5. Konishi, T., J. Hayashi, I. Moriya, and R. Takahashi. 1984. Inheritance and linkage studies in barley. VII. Location of six new mutant genes on chromosome 3. Ber. Ohara Inst. landw. Biol., Okayama Univ. 18:251-264.
6. Lundqvist, U. 1992. Coordinator's report: Eceriferum genes. BGN 21:99-101.
7. Lundqvist, U., and D. von Wettstein. 1962. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. Hereditas 48:342-362.
8. Lundqvist, U., and D. von Wettstein. 1971. Stock list for the eceriferum mutants. BGN 1:97-102.
9. Lundqvist, U., and D. von Wettstein. 1973. Stock list for the eceriferum mutants II. BGN 3:110-112.
10. Lundqvist, U., and D. von Wettstein. 1975. Stock list for the eceriferum mutants III. BGN 5:88-91.
11. Lundqvist, U., and D. von Wettstein. 1977. Stock list for the eceriferum mutants IV. BGN 7:92-96.
12. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.
13. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.
14. Lundqvist, U., P. von Wettstein-Knowles, and D. von Wettstein. 1968. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. II. Hereditas 59:473-504.
15. McProud, W.L., and R.F. Eslick. 1971. Allelism within a series of glossy sheath mutants. BGN 1:31-34.
16. Rasmusson, D.C., and J.W. Lambert. 1965. Inheritance of the glossy-sheath character in barley, Hordeum vulgare L. Crop Sci. 5:251-253.
17. Robertson, D.W., G.A. Wiebe, R.G. Shands, and A. Hagberg. 1965. A summary of linkage studies in cultivated barley, Hordeum species: Supplement III, 1954-1963. Crop Sci. 5:33-43.
18. Singh, R.J., and T. Tsuchiya. 1974. Further information on telotrisomic analysis in barley. BGN 4:66-69.
19. Wettstein-Knowles, P. von. 1991. New alleles of Cer-yy and cer-b. BGN 20:66-68.

W.L. McProud and R.F. Eslick. 1971. BGN 1:169.
U. Lundqvist. 1975. BGN 5:116 as BGS 398, Eceriferum-b, cer-b.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:294-295.

Stock number: BGS 353
Locus name: Glossy sheath 3
Locus symbol: gsh3

Previous nomenclature and gene symbolization:

Glossy sheath 3 = gs3 (20).
Eceriferum-a = cer-a (4, 17).

Monofactorial recessive (4, 14).
Located in chromosome 7HS [1S] (2, 14, 19, 20), about 2.1 cM proximal from the fch12 (chlorina seedling 12) locus (16), and 6.5 to 12.9 cM proximal from the brh1 (brachytic 1) locus (15, 16, 18).

Surface wax coating appears absent on the spike, leaf sheath, and stem (wax code - - ++) (4, 14, 20). Concerning the chemical epicuticular wax composition, only traces of the -diketone wax classes and alkan-2-ol containing esters occur in the cer-a.6 mutant in Bonus (21, 22).

A radiation induced mutant in Mars (CIho 7015) (14, 19, 20); an X-ray induced mutant in Maja (PI 184884) (4).

gsh3.i in Mars (13, 14); gsh3.j in Chevron (CIho 1111), gsh3.k in a hybrid between Atlas and Algerian (13); cer-a.1 in Maja (1, 4, 5); cer-a.6, -a.12 (trans), -a.33 in Bonus (PI 189763) (4, 5); cer-a.98 (trans), -a.121 in Bonus, -a.207, -a.215, -a.224, -a.225, -a.235, -a.242, -a.276, -a.302, -a.329, -a.503 in Foma (CIho 11333) (5, 12); cer-a.134, -a.190, -a.191, -a.192, a.682 in Bonus, -a.224, -a.534, -a.539 in Foma, -a.1007, -a.1013, -a.1021, -a.1028, -a.1050, -a.1051 in Carlsberg II (CIho 10114) (5); cer-a.154, -a.160, -a.750, -a.857, -a.862, -a.863 in Bonus (6); cer-a.836, -a.873, -a.896, -a.916 in Bonus, -a.1194 in Kristina (7); cer-a.1197, -a.1249, -a.1250 in Kristina (8); cer-a.1199, -a.1308, -a.1336, -a.1347 in Kristina (9); cer-a.913, -a.1394, -a.1461, -a.1463 in Bonus, -a.1509, -a.1513 in Nordal (10); cer-a.1524, -a.1530, -a.1531, -a.1554, -a.1557 in Nordal, -a.1704, -a.1570, -a.1774, -a.1788 in Bonus (11); gsh3.af (OUM023), gsh3.aj (OUM027) in Akashinriki (PI 467400, OUJ659) (3).
Mutant used for description and seed stocks:
gsh3.i in Mars (GSHO 737); cer-a.1 in Maja (GSHO 422); gsh3.i in Bowman (PI 483237)*8 (GSHO 1825); cer-a.1 in Maja is used for allelism tests; cer-a.6 in Bonus is used for wax chemistry and wax structure studies.

1. Eslick, R.F. 1974. Allele tests of gs and cer mutants. BGN 4:9-11.
2. Fester, T., and B. Søgaard. 1969. The localization of eceriferum loci in barley. Hereditas 61:327-337.
3. Konishi, T. 1973. Genetic analyses of EMS-induced mutants in barley. BGN 3:28-31.
4. Lundqvist, U., and D. von Wettstein. 1962. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. Hereditas 48:342-362.
5. Lundqvist, U., and D. von Wettstein. 1971. Stock list for the eceriferum mutants. BGN 1:97-102.
6. Lundqvist, U., and D. von Wettstein. 1973. Stock list for the eceriferum mutants II. BGN 3:110-112.
7. Lundqvist, U., and D. von Wettstein. 1975. Stock list for the eceriferum mutants III. BGN 5:88-91.
8. Lundqvist, U., and D. von Wettstein. 1977. Stock list for the eceriferum mutants IV. BGN 7:92-96
9. Lundqvist, U., and D. von Wettstein. 1979. Stock list for the eceriferum mutants V. BGN 9:135-137.
10. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.
11. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.
12. Lundqvist, U., P. von Wettstein-Knowles, and D. von Wettstein. 1968. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. II. Hereditas 59:473-504.
13. McProud, W.L., and R.F. Eslick. 1971. Allelism within a series of glossy sheath mutants. BGN 1:31-34.
14. Rasmusson, D.C., and J. W. Lambert. 1965. Inheritance of the glossy-sheath character in barley, Hordeum vulgare L. Crop Sci. 5:251-253.
15. Shahla, A., and T. Tsuchiya. 1987. Cytogenetic studies in barley chromosome 1 by means of telotrisomic, acrotrisomic and conventional analysis. Theor. Appl. Genet. 75:5-12.
16. Søgaard, B. 1977. The localization of eceriferum loci in barley. V. Three point tests of genes on chromosome 1 and 3 in barley. Carlsberg Res. Commun. 42:67-75.
17. Tsuchiya, T. 1972. Allelism testing of various mutations in barley. BGN 2:80-87.
18.Tsuchiya, T. 1986. Linkage maps of barley (Hordeum vulgare L.). BGN 16:40-43.
19. Tsuchiya, T., and T.E. Haus. 1973. Allelism testing in barley. I. Analysis of ten mapped genes. J. Hered. 64:282-284.
20. Walker, G.W.R., J. Dietrich, R. Miller, and K. Kasha. 1963. Recent barley mutants and their linkages II. Genetic data for further mutants. Can. J. Genet. Cytol. 5:200-219.
21. Wettstein-Knowles, P. von. 1972. Genetic control of -diketone and hydroxy- -diketone synthesis in epicuticular waxes in barley. Planta 106:113-130.
22. Wettstein-Knowles, P. von. 1976. Biosynthetic relationships between -diketones and esterified alkan-2-ols deduced from epicuticular wax of barley mutants. Mol. & Gen. Genet. 144:43-48.

W.L. McProud and R.F. Eslick. 1971. BGN 1:169-170.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:296-297.

Stock number: BGS 354
Locus name: Glossy sheath 4
Locus symbol: gsh4

Previous nomenclature and gene symbolization:

Glossy sheath 4 = gs4 (17).
Eceriferum-x = cer-x (5).

Monofactorial recessive (5, 17).
Located in chromosome 6HL (15, 16, 17), near the centromere, and less than 3.0 cM distal from the rob (orange lemma) locus (1, 2, 3).

Surface wax coating appears absent on the spike, leaf sheath, and stem (wax code - - ++) (5, 15). Concerning the chemical epicuticular wax composition, only traces of the -diketone wax classes and alkan-2-ol containing esters occur in the cer-x mutant in Bonus (18, 19).

A radiation induced mutant in Gateway (CIho 10072) (14, 15); an ethylene imine induced mutant in Bonus (PI 189763) (5).

gsh4.l Gateway (14, 15); cer-x.60 in Bonus (4, 5, 6, 14); cer-x.102, -x.114 in Bonus, -x.228 (GSHO 445), -x.309, -x.350, -x.417, -x.449, -x.450, -x.507 in Foma (CIho 11333) (6, 13); cer-x.164, -x.658 in Bonus, -x.561 in Foma, -x.1049, -x.1055, -x.1066 in Carlsberg II (CIho 10114) (6); cer-x.788 in Bonus, -x.1095 in Kristina (7); cer-x.1180 in Kristina (8); cer-x.1236 in Kristina (9); cer-x.886 in Bonus (10); cer-x.1398, -x.1435 in Bonus, -x.1508 in Nordal (11); cer-x.1000, -x.1498 in Bonus, -x.1518, -x.1523, -x.1529, -x.1536, -x.1547, -x.1552 in Nordal, -x.1732, -x.1756 in Bonus (12).
Mutant used for description and seed stocks:
gsh4.l in Gateway (GSHO 738); cer-x.60 in Bonus (GSHO 1520); gsh4.l in Bowman (PI 483237)*7 (GSHO 2072); cer-x.60 in Bowman*7 (GSHO 2073).

1. Eslick, R.F., R.T. Ramage, and D.R. Clark. 1974. Two genetic male steriles, msg6 and msg,,bk, assigned to chromosome 6. BGN 4:11-15.
2. Falk, D.E., M.J. Swartz, and K.J. Kasha. 1980. Linkage data with genes near the centromere of barley chromosome 6. BGN 10:13-16.
3. Kasha, K.J., D.E. Falk, and A. Ho-Tsai. 1978. Linkage data with genes on chromosome 6. BGN 8:61-65.
4. Lundqvist, U. 1992. Coordinator's report: Eceriferum genes. BGN 21:99-101.
5. Lundqvist, U., and D. von Wettstein. 1962. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. Hereditas 48:342-362.
6. Lundqvist, U., and D. von Wettstein. 1971. Stock list for the eceriferum mutants. BGN 1:97-102.
7. Lundqvist, U., and D. von Wettstein. 1973. Stock list for the eceriferum mutants II. BGN 3:110-112.
8. Lundqvist, U., and D. von Wettstein. 1975. Stock list for the eceriferum mutants III. BGN 5:88-91.
9. Lundqvist, U., and D. von Wettstein. 1977. Stock list for the eceriferum mutants IV. BGN 7:92-96.
10. Lundqvist, U., and D. von Wettstein. 1979. Stock list for the eceriferum mutants V. BGN 9:135-137.
11. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.
12. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.
13. Lundqvist, U., P. von Wettstein-Knowles, and D. von Wettstein. 1968. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. II. Hereditas 59:473-504.
14. McProud, W.L., and R.F. Eslick. 1971. Allelism within a series of glossy sheath mutants. BGN 1:31-34.
15. Rasmusson, D.C., and J.W. Lambert. 1965. Inheritance of the glossy-sheath character in barley, Hordeum vulgare L. Crop Sci. 5:251-253.
16. Shahla, A., and T. Tsuchiya. 1990. Genetic analysis in six telotrisomic lines in barley (Hordeum vulgare L.). J. Hered. 81:127-130.
17. Walker, G.W.R., J. Dietrich, R. Miller, and K. Kasha. 1963. Recent barley mutants and their linkages II. Genetic data for further mutants. Can. J. Genet. Cytol. 5:200-219.
18. Wettstein-Knowles, P. von. 1972. Genetic control of -diketone and hydroxy- -diketone synthesis in epicuticular waxes in barley. Planta 106: 113-130.
19. Wettstein-Knowles, P. von. 1976. Biosynthetic relationships between -diketones and esterified alkan-2-ols deduced from epicuticular wax of barley mutants. Mol. & Gen. Genet. 144:43-48.

W.L. McProud and R.F. Eslick. 1971. BGN 1:170-171.
U. Lundqvist. 1975. BGN 5:134 as BGS 416, Eceriferum-x, cer-x.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:298-299.

Stock number: BGS 355
Locus name: Glossy sheath 5
Locus symbol: gsh5

Previous nomenclature and gene symbolization:

Glossy sheath 5 = gs5 (12).
Eceriferum-s = cer-s (2, 10).
Glossy sheath 8 = gs8 (14).

Monofactorial recessive (2).
Located in chromosome 2HL (12, 13, 14, 16); gsh5 is in the centromeric region, about 2.5 cM proximal from the eog (elongated outer glume) locus, and about 32.0 cM proximal from the vrs1 (six-rowed spike 1) locus (12, 14).

Mutants have a reduced surface wax covering on the spikes and stems, and a bright glossy green color on all leaf sheaths (12) (wax code + - ++) (2). The nodes appear to have excessive wax covering. The cer-s.31 allele is associated with delayed heading in the Bowman backcross-derived line, but the gsh5.m allele is not.

An X-ray induced mutant in Bonus (PI 189763) (2); a radiation induced mutant in Jotun (PI 467357) (12); a spontaneous mutant in Okaiku 3 (OUL011) (15).

cer-s.24 in Bonus (2); cer-s.31 in Bonus (1, 2); cer-s.41 in Bonus (2); cer-s.287, -s.303, -s.372, -s.444 in Foma (CIho 11333) (10); cer-s.1047 in Carlsberg II (CIho 10114) (3); cer-s.137 in Bonus, -s.591 in Foma, -s.803 in Bonus, -s.1132 in Kristina (4); cer-s.914 in Bonus (5); cer-s.622 in Bonus, -s.1054 in Carlsberg II, -s.1153 in Kristina (6); cer-s.1323 in Bonus (7); cer-s.1134 in Kristina (8); cer-s.1754 in Bonus (9); gsh5.m in Jotun (12); gsh5.n in Okaiku 3 (GSHO 832) (11, 15).
Mutant used for description and seed stocks:
gsh5.m in Jotun (GSHO 739); cer-s.31 in Bonus (GSHO 440); cer-s.31 in Bowman (PI 483237)*5 (GSHO 1888); gsh5.m in Bowman*8 (GSHO 1889).

1. Eslick, R.F. 1974. Allele tests of gs and cer mutants. BGN 4:9-11.
2. Lundqvist, U., and D. von Wettstein. 1962. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. Hereditas 48:342-362.
3. Lundqvist, U., and D. von Wettstein. 1971. Stock list for the eceriferum mutants. BGN 1:97-102.
4. Lundqvist, U., and D. von Wettstein. 1973. Stock list for the eceriferum mutants II. BGN 3:110-112.
5. Lundqvist, U., and D. von Wettstein. 1975. Stock list for the eceriferum mutants III. BGN 5:88-91.
6. Lundqvist, U., and D. von Wettstein. 1977. Stock list for the eceriferum mutants IV. BGN 7:92-96.
7. Lundqvist, U., and D. von Wettstein. 1979. Stock list for the eceriferum mutants V. BGN 9:135-137.
8. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.
9. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.
10. Lundqvist, U., P. von Wettstein-Knowles, and D. von Wettstein. 1968. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. II. Hereditas 59:473-504.
11. McProud, W.L., and R.F. Eslick. 1971. Allelism within a series of glossy sheath mutants. BGN 1:31-34.
12. Rasmusson, D.C., and J.W. Lambert. 1965. Inheritance of the glossy-sheath character in barley, Hordeum vulgare L. Crop Sci. 5:251-253.
13. Singh, R.J., and T. Tsuchiya. 1974. Further information on telotrisomic analysis in barley. BGN 4:66-69.
14. Takahashi, R., J. Hayashi, T. Konishi, and I. Moriya. 1972. Inheritance and linkage studies in barley. V. Locating seven new mutant genes. Ber. Ohara Inst. landw. Biol., Okayama Univ. 15:147-168.
15. Takahashi, R., J. Hayashi, and I. Moriya. 1971. Linkage studies in barley. BGN 1:51-58.
16. Tsuchiya, T. 1972. Cytogenetics of telotrisomics in barley. BGN 2:93-98.

W.L. McProud and R.F. Eslick. 1971. BGN 1:171-172.

J.D. Franckowiak and U. Lundqvist. 1997. BGN 26:300-301.

Stock number: BGS 356
Locus name: Glossy sheath 6
Locus symbol: gsh6

Previous nomenclature and gene symbolization:

Glossy sheath 6 = gs6 (15).
Eceriferum-c = cer-c (4).
Eceriferum-cqu = cer-cqu (13, 21).

Monofactorial recessive (4, 15, 17).
Located in chromosome 2HS (17, 18, 19), in a subterminal position (16), and over 33.7 cM distal from the Eam1 (early maturity 1) locus (3).

A bright, glossy green color is apparent on the spike, leaf sheath, and stem (15). The gsh6 locus is the cer-c component of the complex locus cer-cqu (1), which controls formation of -diketone lipids in epicuticular wax layer (21, 22). Most alleles have the wax code - - ++, but a few alleles show delayed development of surface waxes on the sheath (wax code - +/- ++) (4, 14).

An X-ray induced mutant in Bonus (PI 189763) (4), a radiation induced mutant in Domen (CIho 9562) (15).

cer-c.3 in Bonus (1, 4); cer-c.7, -c.29, -c.36, -c.44, -c.61, -c.63 in Bonus (4, 5); cer-c.73, -c.89, -c.95, -cu.108, -c.115, -c.123, -cqu.124 in Bonus, -c.201, -c.206, -c.220, -c.223, -c.234, -c.238, -c.240, -c.250, -c.270, -c.282, -c.283, -c.284, -c.290, -c.291, -c.293. -c.306, -c.315, -c.332, -c.343, -c.354, -c.361, -c.364, -c.367, -c.368, -c.374, -c.379, -c.398h, -c.398l, -c.405, -c.406, -c.413, -cqu.416, -cqu.420, -c.431, -c.445, -c.455, -c.469, -c.471, -c.502 in Foma (CIho 11333) (5, 13); cer-c.143, -c.171, -c.175 in Bonus, -c.345, -c.451, -c.459, -c.508, -c.513, -c.514, -c.520, -c.526, -c.544, -c.550, -c.563, -c.596, -c.601, -c.661 in Foma, -c.669, -c.673, -c.690, -c.693, -c.698, -c.703, -c.711, -cqu.724, -cqu.733 in Bonus, -c.1001, -c.1004, -c.1008, -c.1010, -c.1015, -c.1022, -c.1029, -c.1034, -c.1035, -c.1039, -c.1062, -c.1067, -c.1082 in Carlsberg II (CIho 10114) (5); cer-c.589 in Foma, -c.748, -c.760, -c.764, -c.767, -c.770, -c.774, -c.786, -c.793, -c.794, -c.841, -c.845, -c.847, -c.851, -c.860, -c.870, -c.881, -c.883 in Bonus, -c.1032 in Carlsberg II, -c.1098, -c.1100, -c.1109, -c.1117, -c.1123, -c.1125, -c.1135, -c.1136, -c.1140, -c.1147 in Kristina (6); cer-c.499, -c.609 in Foma, -c.630, -c.635, -c.640, -cu.644, -cq.645, -c.648, -c.890, -c.897, -c.902, -c.903, -c.904, -c.907, -c.912, -c.915, -cqu.925, -c.942, -cqu.944, -c.945, -c.980 in Bonus, -c.1156, -c.1187 in Kristina (7); cer-c.144, -cu.947, -c.987 in Bonus, -c.1110, -c.1212, -c.1229, -c.1248, -c.1258, -c.1264, -c.1276 in Kristina (8); cer-c.511 in Foma, -c.1200, -c.1254, -c.1278, -c.1287, -c.1291 in Kristina, -c.1319, -c.1325, -c.1329, -c.1334, -c.1338 in Bonus (9); cer-c.993, -c.994, -c.995, -c.999, -c.1348, -c.1355, -c.1359, -c.1365, -c.1376, -c.1381, -c.1382, -c.1385, -c.1386, -c.1389, -c.1416, -c.1421, -c.1439, -c.1448, -c.1460, -c.1462, -c.1466, -c.1467 in Bonus, -c.1501, -c.1502, -c.1503, -c.1505, -c.1506, -c.1511, -c.1515 in Nordal (10); cer-c.1491, -c.1701, -c.1702, -c.1703, -c.1705, -c.1707, -c.1714, -c.1737, -c.1739, -c.1749, -c.1755, -c.1757 in Bonus, -c.1516, -c.1520, -c.1525, -c.1526, -c.1532, -c.1553, -c.1555, -c.1560 in Nordal (11); cer-c.602a in Foma (12); gsh6.o in Domen, gsh6.p in PI 184877, gsh6.q in Kogane Mugi (OUL010, PI 225020), gsh6.r in Barbless (CIho 5105), gsh6.s in Betzes (PI 129430), gsh6.t in Heines Hanna (PI 539131), gsh6.u, gsh6.v, gsh6.w, and gsh6.x in Compana (PI 539111) (14); gsh6.z (OUM017), gsh6.ac (OUM020), gsh6.ai (OUM026) in Akashinriki (PI 467400, OUJ659) (2); gsh6.aq in an Ethiopian introduction (PI 151787, OUE 120) (2).

gsh6.s in Betzes (GSHO 740); cer-c.3 in Bonus (GSHO 212, GSHO 424, GSHO 959); gsh6.s in Bowman (PI 483237)*7 (GSHO 1869).

1. Eslick, R.F. 1974. Allele tests of gs and cer mutants. BGN 4:9-11.
2. Konishi, T. 1973. Genetic analyses of EMS-induced mutants in barley. BGN 3:28-31.
3. Luna Villafaña, A. 1995. Mapping marker genes in chromosome 2 of barley. M.S. Thesis. North Dakota State Univ., Fargo.
4. Lundqvist, U., and D. von Wettstein. 1962. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. Hereditas 48:342-362.
5. Lundqvist, U., and D. von Wettstein. 1971. Stock list for the eceriferum mutants. BGN 1:97-102.
6. Lundqvist, U., and D. von Wettstein. 1973. Stock list for the eceriferum mutants II. BGN 3:110-112.
7. Lundqvist, U., and D. von Wettstein. 1975. Stock list for the eceriferum mutants III. BGN 5:88-91.
8. Lundqvist, U., and D. von Wettstein. 1977. Stock list for the eceriferum mutants IV. BGN 7:92-96.
9. Lundqvist, U., and D. von Wettstein. 1979. Stock list for the eceriferum mutants V. BGN 9:135-137.
10. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.
11. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.
12. Lundqvist, U., and D. von Wettstein. 1988. Stock list for the eceriferum mutants VIII. BGN 18:88-91.
13. Lundqvist, U., P. von Wettstein-Knowles, and D. von Wettstein. 1968. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. II. Hereditas 59:473-504.
14. McProud, W.L., and R.F. Eslick. 1971. Allelism within a series of glossy sheath mutants. BGN 1:31-34.
15. Rasmusson, D.C., and J.W. Lambert. 1965. Inheritance of the glossy-sheath character in barley, Hordeum vulgare L. Crop Sci. 5:251-253.
16. Schondelmaier, J., G. Fischbeck, and A. Jahoor. 1993. Linkage studies between morphological and RFLP markers in the barley genome. BGN 22:57-62.
17. Singh, R.J., and T. Tsuchiya. 1974. Further information on telotrisomic analysis in barley. BGN 4:66-69.
18. Takahashi, R., J. Yamamoto, S. Yasuda, and Y. Itano. 1953. Inheritance and linkage studies in barley. Ber. Ohara Inst. landw. Forsch. 10:29-52.
19. Tsuchiya, T. 1972. Cytogenetics of telotrisomics in barley. BGN 2:93-98.
20. Wettstein-Knowles, P. von. 1972. Genetic control of -diketone and hydro- -diketone synthesis in epicuticular waxes of barley. Planta 106:113-130.
21. Wettstein-Knowles, P. von. 1992. Molecular genetics of the lipid synthesis in barley. p. 753-771. In L. Munck, K. Kirkegaard, and B. Jensen (eds.) Barley Genetics VI, Vol. II. Proc. Sixth Int. Barley Genet. Symp., Helsingborg, Sweden, 1991. Munksgaard International Publishers Ltd., Copenhagen.

W. L. McProud and R.F. Eslick. 1971. BGN 1:172-173.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:302-303.

Stock number: BGS 357
Locus name: Male sterile genetic 1
Locus symbol: msg1

Previous nomenclature and gene symbolization:

Male sterile = ms (12).
Male sterile 1 = ms1 (3).

Monofactorial recessive (14).
Located in chromosome 1HL [5L] (11), near the centromere (6), and about 10.0 cM proximal from the nec1 (necrotic leaf spot 1) locus (9, 12).

Selfing - none (5).
Outcrossing - complete female fertility (5).
Stamens - anthers smaller than fertile sib, no stomium or filament elongation (13).
Pollen - microspores degenerate at or before the free microspore stage (10); non-staining, shrunken, and devoid of cytoplasm (13).
Cytology - normal development and differentiation of anthers until completion of meiosis (10).

A spontaneous mutant in the Composite Cross line CIho 5368 (14).

msg1.ca in CIho 5368 (MSS005) (10, 14); msg1.i (MSS077) in 80TT25 (CIho 13638), msg1.t (MSS042) in Trophy (CIho 10647), msg1.ai (MSS100) in Betzes (PI 129430) (7, 8); msg1.ar (MSS310) in Glossy Brachytic (CIho 15246), msg1.bp (MSS330) in Betzes (1, 8); msg1.cz (MSS348) in Betzes (2, 8); msg1.gb (MSS429) in Maris Mink (3); msg1.jv (MSS527) in a Harrington outcross (4).

msg1.ca in Betzes*11(GSHO 1810); msg1.ca in Bowman (PI 483237)*8 (GSHO 2042).

1. Hockett, E.A. 1973. The genetic male sterile collection. BGN 3:87-89.
2. Hockett, E.A. 1977. The genetic male sterile collection. BGN 7:97-100.
3. Hockett, E.A. 1984. Coordinator's report. The genetic male sterile barley collection. BGN 14:70-75.
4. Hockett, E.A. 1988. New mutants in the genetic male sterile collection. BGN 18:70-73.
5. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
6. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
7. Hockett, E.A., R.F. Eslick, D.A. Reid, and G.A. Wiebe. 1968. Genetic male sterility in barley. II. Available spring and winter stocks. Crop Sci. 8:754-755.
8. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
9. Jensen, J., and J.H. Jørgensen. 1973. Locating some genes on barley chromosome 5. BGN 3:25-27.
10. Kaul, C.L., and S.P. Singh. 1966. Studies in male-sterile barley. II. Pollen abortion. Crop Sci 6:539-541.
11. Ramage, R.T., C.R. Burnham, and A Hagberg. 1961. A summary of translocation studies in barley. Crop Sci. 1:277-279.
12. Ramage, T., and J.L.A. Eckhoff. 1985. Assignment of mutants in Morex to chromosomes. BGN 15:22-25.
13. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.
14. Suneson, C.A. 1940. A male sterile character in barley. A new tool for the plant breeder. J. Hered. 31:213-214.

E.A. Hockett. 1971. BGN 1:175.

J.D. Franckowiak and U. Lundqvist. 1997. BGN 26:304-305.

Stock number: BGS 358
Locus name: Male sterile genetic 2
Locus symbol: msg2

Previous nomenclature and gene symbolization:

Male sterile 2 = ms₂ (1).
Male sterile 2 = ms2 (3).

Monofactorial recessive (1).
Located in chromosome 2HL (1, 9), about 2.4 cM proximal from the eog (elongated outer glume) locus (6).

Selfing - none (3).
Outcrossing - complete female fertility (3).
Stamens - anthers smaller than fertile sib, no stomium or filament elongation (8).
Pollen - non-staining, shrunken, and no normal grains (8).

A spontaneous mutant in the F₂ progeny of Manchuria (CIho 2330) X CIho 4363 (3).

msg2.cb (MSS046) in F₂ of Manchuria/CIho 4363 (3, 5); msg2.ax (MSS045) in Compana (PI 539111) (4, 5); msg2.ed (MSS379) in Ingrid (CIho 10083) (2, 7).

msg2.cb in Herta*10 (GSHO 2371), msg2.cb in Manchuria*19 (MSS047), msg2.cb in Ogalitsu*14 (MSS048), msg2.cb in Trebi*19 (MSS050) (4); msg2.cb from Herta*10 in Bowman (PI 483237)*7 (GSHO 1890).

1. Austenson, H.M. 1948. Linkage relations of the male sterile gene ms₂ in barley. M.S. Thesis. Univ. of Saskatchewan, Saskatoon.
2. Hockett, E.A. 1984. Coordinator's report. The genetic male sterile barley collection. BGN 14:70-75.
3. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
4. Hockett, E.A., R.F. Eslick, D.A. Reid, and G.A. Wiebe. 1968. Genetic male sterility in barley II. Available spring and winter stocks. Crop Sci. 8:754-755.
5. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
6. Jin, Y., J.D. Franckowiak, and G.D. Statler. 1993. Linkage among some of the morphological markers in Wolfe's stocks. BGN 22:25-26.
7. Ramage, R.T., and J.E. Flora. 1981. Allele tests and chromosome location of two male sterile mutants in Ingrid. BGN 11:36-37.
8. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.
9. Tsuchiya, T., and R.J. Singh. 1973. Further information on telotrisomics analysis in barley. BGN 3:75-79.

E.A. Hockett. 1971. BGN 1:175-176.

J.D. Franckowiak. 1997. BGN 26:306.

Stock number: BGS 359
Locus name: Male sterile genetic 3
Locus symbol: msg3

Previous nomenclature and gene symbolization:

Male sterile 10 = ms10 (1, 5).
Male sterile 3 = ms3 (1, 3).

Monofactorial recessive (3).
Located in chromosome 2HS (3), about 4.6 cM distal from the eog (elongated outer glume) locus, and 0.6 cM distal from the fch1 (chlorina seedling 1) locus (3).

Selfing - none (1, 3).
Outcrossing - complete female fertility (1, 3).
Stamens - anthers much smaller than fertile sib (1), no stomium or filament elongation (4).
Pollen - non-staining, no free pollen grains (4).
The male sterile plants are also about 1/3 normal size with short, wide leaves (1, 3). Spikes are dense and spikelets are small and malformed, awns are 1/3 normal length, and double or triple pistils (fasciation) occur in some spikelets (3).

An acetone induced mutant in Gateway (CIho 10072) (3).

msg3.cc in Gateway (MSS051) (2, 3).

msg3.cc in Gateway (GSHO 1130); msg3.cc in Bowman (PI 483237)*7 (GSHO 1885).

1. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
2. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
3. Kasha, K.J., and G.W.R. Walker. 1960. Several recent barley mutants and their linkages. Can. J. Genet. Cytol. 2:397-415.
4. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.
5. Robertson, D.W., G.A. Wiebe, R.G. Shands, and A. Hagberg. 1965. A summary of linkage studies in cultivated barley, Hordeum species: Supplement III. 1954-1963. Crop Sci. 5:33-43.

E.A. Hockett. 1971. BGN 1:176.

J. D. Franckowiak. 1997. BGN 26:307.

Stock number: BGS 360
Locus name: Male sterile genetic 4
Locus symbol: msg4

Previous nomenclature and gene symbolization:

Male sterile 4 = ms4 (1).

Monofactorial recessive (1).
Located in chromosome 1H [5], near the centromere (2).

Selfing - none (1).
Outcrossing - complete female fertility (1).
Stamens - anthers smaller than fertile sib, no stomium or filament elongation (4).
Pollen - non-staining, shrunken, and no normal grains (4).

A spontaneous mutant in Freja (CIho 7130) (1).

msg4.cd in Freja (MSS052) (1, 3).

msg4.cd in Freja (GSHO 2392); msg4.cd in Bowman (PI 483237)*7 (GSHO 2043).

1. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
2. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
3. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
4. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.

E.A. Hockett. 1971. BGN 1:177-178.

J.D. Franckowiak. 1997. BGN 26:308.

Stock number: BGS 361
Locus name: Male sterile genetic 5
Locus symbol: msg5

Previous nomenclature and gene symbolization:

Male sterile 5 = ms5 (4).

Monofactorial recessive (4).
Located in chromosome 3HS (5), about 13.1 cM proximal from the uzu (uzu) locus, and about 6.8 cM proximal from the alm (albino lemma) locus (1).

Selfing - none (4).
Outcrossing - complete female fertility (4).
Stamens - anthers smaller than fertile sib, no stomium or filament elongation (7).
Pollen - non-staining, some normal-appearing grains (7).

A spontaneous mutant in Carlsberg II (CIho 10114) (4).

msg5.ce in Carlsberg II (MSS053) (4, 6); msg5.s (MSS087) in Schweigers Erika (CIho 11501) (2, 6); msg5.fr (MSS419) in Midas (PI 343078) (3).

msg5.ce in Carlsberg II (GSHO 2403); msg5.ce in Bowman (PI 483237)*7 (GSHO 1954).

1. Eslick, R.F., W.L. McProud. 1974. Positioning of the male sterile 5 (msg5) on chromosome 3. BGN 4:16-23.
2. Hockett, E.A. 1972. Coordinator's report on the genetic male sterile barley collection. BGN 2:139-144.
3. Hockett, E.A. 1984. Coordinator's report. The genetic male sterile barley collection. BGN 14:70-75.
4. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
5. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
6. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
7. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.

E.A. Hockett. 1971. BGN 1:178.

J.D. Franckowiak. 1997. BGN 26:309.

Stock number: BGS 362
Locus name: Male sterile genetic 6
Locus symbol: msg6

Previous nomenclature and gene symbolization:

Male sterile 6 = ms6 (4).

Monofactorial recessive (4).
Located in chromosome 6HS (3), near the centromere (2, 6), and about 1.0 cM distal from the rob (orange lemma) locus (2).

Selfing - none (2), but occasionally a few selfed seeds occur.
Outcrossing - complete female fertility (2).
Stamens - equal in size to normal anthers (2); stomium present, filament elongation (7).
Pollen - nearly normal stained in both field and greenhouse plants, most grains appear normal (7); grains are non-functional because aperture development is abnormal (1).
Cytology - normal development and differentiation of anthers (7, 8).

A spontaneous mutant in Heines Hanna (PI 539131) (4, 5).

msg6.cf in Heines Hanna (MSS054) (4, 5).

msg6.cf in Heines Hanna (GSHO 2405); msg6.cf in Bowman (PI 483237)*7 (GSHO 2078).

1. Ahokas, H. 1975. Male sterile mutants of barley. I. Inaperturate pollen of the msg6cf mutant. Ann. Bot. Fenn. 12:17-21.
2. Eslick, R.F., R.T. Ramage, and D.R. Clark. 1974. Two genetic male steriles, msg6 and msg,,bk, assigned to chromosome 6. BGN 4:11-15.
3. Falk, D.E. 1994. Creation of a marked telo 6S trisomic for chromosome 6. BGN 23:32.
4. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
5. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
6. Lehmann, L., and P. Hagberg. 1978. Linkage studies of msg6 using four translocations involving chromosomes 5 and 6. BGN 8:73-74.
7. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.
8. Roath, W.W., and E.A. Hockett. 1971. Pollen development in genetic male-sterile barley. p. 308-315. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.

E.A. Hockett. 1971. BGN 1:178-179.

J.D. Franckowiak. 1997. BGN 26:310.

Stock number: BGS 363
Locus name: Male sterile genetic 7
Locus symbol: msg7

Previous nomenclature and gene symbolization:

Male sterile 7 = ms7 (5).

Monofactorial recessive (5).
Located in chromosome 5HL [7L], about 5.5 cM from the raw1 (smooth awn 1) locus (1).

Selfing - none (5).
Outcrossing - complete female fertility (5).
Stamens - anthers much smaller than fertile sibs, no stomium or filament elongation (7).
Pollen - microspores degenerate before the free microspore stage, non-staining, no free grains (7, 8).
Cytology - normal development and differentiation of anthers until completion of meiosis, but degeneration of tapetal tissue at the tetrad stage (8).

A spontaneous mutant in Dekap (CIho 3351) (4).

msg7.cg in Dekap (MSS055) (2, 4); msg7.ah (MSS099) in HB 421/78 (CIho 13641) (2, 6); msg7.fx (MSS425) in Proctor (3, 4).

msg7.cg in Dekap (GSHO 2406); msg7.cg in Bowman (PI 483237)*7 (GSHO 2109).

1. Franckowiak, J.D. 1991. Association of male sterility genes with a specific chromosome using multiple marker stocks. BGN 20:31-36.
2. Hockett, E.A. 1976. The genetic male sterile collection. BGN 6:108.
3. Hockett, E.A. 1984. Coordinator's report. The genetic male sterile barley collection. BGN 14:70-75.
4. Hockett, E.A. 1991. The identification of eight new loci and allelism of 14 additional mutants. BGN 20:37-40.
5. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
6. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
7. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.
8. Roath, W.W., and E.A. Hockett. 1971. Pollen development in genetic male-sterile barley. p. 308-315. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.

E.A. Hockett. 1971. BGN 1:179-180.

J.D. Franckowiak. 1997. BGN 26:311.

Stock number: BGS 364
Locus name: Male sterile genetic 8
Locus symbol: msg8

Previous nomenclature and gene symbolization:

Male sterile 8 = ms8 (4).

Monofactorial recessive (4).
Located in chromosome 5HL [7L] (2, 5), about 8.3 cM from the raw1 (smooth awn 1) locus (2).

Selfing - none (4).
Outcrossing - complete female fertility (4).
Stamens - anthers smaller than fertile sibs (4), stomium present, and filament elongation (1, 7).
Pollen - reduced staining (6.9% stains with 2,3,5-triphenyltetrazolium, 20.7% stains with acetocarmine), some grains appear normal (7, 8).
Cytology - normal development and differentiation of microspores up to the free microspore stage (8). Grains are undeveloped, have a stainable ring at the apertural annulus without any actual pore; the exine staining with Fast Blue B is darker than in normal grains (1).

A spontaneous mutant in Betzes (PI 129430) (4).

msg8.ch in Betzes (MSS056) (4, 6); msg8.au (MSS313) in OAC 21 (CIho 1470) (3, 6).

msg8.ch in Betzes (GSHO 2407); msg8.ch in Bowman (PI 483237)*6 (GSHO 2110).

1. Ahokas, H. 1976. Male sterile mutants of barley. III. Additional inaperturate mutants. BGN 6:4-6.
2. Franckowiak, J.D. 1991. Association of male sterility genes with a specific chromosome using multiple marker stocks. BGN 20:31-36.
3. Hockett, E.A. 1975. The genetic male sterile collection. BGN 5:84-86.
4. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
5. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
6. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
7. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.
8. Roath, W.W., and E.A. Hockett. 1971. Pollen development in genetic male-sterile barley. p. 308-315. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.

E.A. Hockett. 1971. BGN 1:180.

J.D. Franckowiak. 1997. BGN 26:312.

Stock number: BGS 365
Locus name: Male sterile genetic 9
Locus symbol: msg9

Previous nomenclature and gene symbolization:

Male sterile 9 = ms9 (2).

Monofactorial recessive (2).
Located in chromosome 2HS (1), over 15.1 cM distal from the eog (elongated outer glume) locus (3).

Selfing - 10% at Bozeman, Montana and 24% at Tucson, Arizona, USA (2).
Outcrossing - complete female fertility (2).
Stamens - anthers smaller than fertile sib (2), stomium present and filament elongation occurs (4).
Pollen - stained in plants grown in the field and in the greenhouse, in some samples the grains appear normal (4).

A spontaneous mutant in Vantage (CIho 7324) (2).

msg9.ci in Vantage (MSS057) (2).

msg9.ci in Vantage (GSHO 2408); msg9.ci in Bowman (PI 483237)*7 (GSHO 1883).

1. Franckowiak, J.D. 1991. Association of male sterility genes with a specific chromosome using multiple marker stocks. BGN 20:31-36.
2. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
3. Luna Villafaña, A. 1995. Mapping marker genes in chromosome 2 of barley. M.S. Thesis. North Dakota State Univ., Fargo.
4. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.

E.A. Hockett. 1971. BGN 1:181.

J.D. Franckowiak. 1997. BGN 26:313.

Stock number: BGS 366
Locus name: Male sterile genetic 10
Locus symbol: msg10

Previous nomenclature and gene symbolization:

Male sterile 3 = ms3 (5).
Male sterile 10 = ms10 (3).

Monofactorial recessive (3).
Located in chromosome 7HS [1S] (1, 4), about 7.2 cM distal from the nud (naked caryopsis) locus (2, 7), and about 14.1 cM from the lks2 (short awn 2) locus (2).

Selfing - none (3).
Outcrossing - complete female fertility (3).
Stamens - anthers smaller than fertile sib, no stomium or filament elongation (8).
Pollen - no staining with 2,3,5-triphenyltetrazolium, but staining with acetocarmine, some normal-appearing grains (8).

A spontaneous mutant in Compana (PI 539111) (5).

msg10.ay in Compana (MSS058) (5, 6); msg10.cy (MSS059) in Manchuria (CIho 2330)*9 (3, 6).

msg10.ay in Compana (GSHO 1811); msg10.ay in Bowman (PI 483237)*7 (GSHO 1835).

1. Eslick, R.F. 1971. Balanced male steriles and dominant pre-flowering selective genes for use in hybrid seed production. p. 292-297. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
2. Eslick, R.F., and E.A. Hockett. 1972. Recombination values of four genes on chromosome 1. BGN 2:123-126.
3. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
4. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
5. Hockett, E.A., R.F. Eslick, D.A. Reid, and G.A. Wiebe. 1968. Genetic male sterility in barley. II. Available spring and winter stocks. Crop Sci. 8:754-755.
6. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
7. Jarvi, A.J., and R.F. Eslick. 1975. Shrunken endosperm mutants in barley. Crop Sci. 15:363-366.
8. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.

E.A. Hockett. 1971. BGN 1:181-182.

J.D. Franckowiak. 1997. BGN 26:314.

Stock number: BGS 367
Locus name: Male sterile genetic 11
Locus symbol: msg11

Previous nomenclature and gene symbolization:

Male sterile 10 = ms10 (2, 3, 4, 5, 7, 8).
Male sterile 11 = ms11 (2, 9).
Male sterile 12 = msg12 (2).

Monofactorial recessive (2, 9).
Location is unknown, but a weak linkage to the raw1 (smooth awn 1) locus is reported (1, 9).

Selfing - none (2).
Outcrossing - complete female fertility (2).
Stamens - anthers smaller than fertile sib, no stomium or filament elongation (6).
Pollen - no staining in msg11.ck plants, but very low staining in msg11.az plants, no normal-appearing grains (6).

A spontaneous mutant in Gateway (CIho 10072) (5).

msg11.ck in Gateway (MSS060) (4, 5); msg11.az (MSS061) in Svalöf 50-109 (CIho 10524) (2, 4).

msg11.az in Svalöf 50-109 (GSHO 1812); msg11.az in Bowman (PI 483237)*7 (GSHO 2299).

1. Franckowiak, J.D. 1991. Association of male sterility genes with a specific chromosome using multiple marker stocks. BGN 20:31-36.
2. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
3. Hockett, E.A., R.F. Eslick, D.A. Reid, and G.A. Wiebe. 1968. Genetic male sterility in barley. II. Available spring and winter stocks. Crop Sci. 8:754-755.
4. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
5. Kasha, K.J., and G.W.R. Walker. 1960. Several recent barley mutants and their linkages. Can. J. Genet. Cytol. 2:397-415.
6. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.
7. Robertson, D.W., G.A. Wiebe, R.G. Shands, and A. Hagberg. 1965. A summary of linkage studies in cultivated barley, Hordeum species: Supplement III. 1954-1963. Crop Sci. 5:33-43.
8. Walker, G.W.R., J. Dietrich, R. Miller, and K. Kasha. 1963. Recent barley mutants and their linkages II. Genetic data for further mutants. Can. J. Genet. Cytol. 5:200-219.
9. Walker, G.W.R., K. Kasha, and R.A. Miller. 1958. Recombination studies in barley. Proc. Genet. Soc. Can. 3:41-43.

E.A. Hockett. 1971. BGN 1:182-183.

J.D. Franckowiak. 1997. BGN 26:315.

Stock number: BGS 368
Locus name: Male sterile genetic 13
Locus symbol: msg13

Previous nomenclature and gene symbolization:

Male sterile 13 = ms13 (1).

Monofactorial recessive (1).
Location is unknown.

Selfing - none (1).
Outcrossing - complete female fertility (1).
Stamens - anthers smaller than those of fertile sibs, no stomium or filament elongation (3).
Pollen - no staining, shrunken, no normal-appearing grains (3).

A spontaneous mutant in Haisa II (CIho 10420) (1).

msg13.cl in Haisa II (MSS062) (1, 2).

msg13.cl in Haisa II (GSHO 1813); msg13.cl in Bowman (PI 483237)*7 (GSHO 2300).

1. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
2. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
3. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.

E.A. Hockett. 1971. BGN 1:183.

J.D. Franckowiak. 1997. BGN 26:316.

Stock number: BGS 369
Locus name: Male sterile genetic 14
Locus symbol: msg14

Previous nomenclature and gene symbolization:

Male sterile 14 = ms14 (5).

Monofactorial recessive (5).
Located in chromosome 7HS [1S] (1, 6), about 2.0 cM from the msg10 (male sterile genetic 10) locus (2).

Selfing - none (5).
Outcrossing - complete female fertility (5).
Stamens - anthers smaller than fertile sib, no stomium or filament elongation (8).
Pollen - less than 1% staining with 2,3,5-triphenyltetrazolium, about 9% with acetocarmine, some normal-appearing grains (8).

A spontaneous mutant in Unitan (CIho 10421) (5).

msg14.cm in Unitan (MSS063) (5, 7); msg14.da (MSS349) in Betzes (PI 129430) (3, 7); msg14.dl (MSS360) in Hector (CIho 15514) (4, 7).

msg14.cm in Unitan (GSHO 1814); msg14.cm in Bowman (PI 483237)*7 (GSHO 1836).

1. Eslick, R.F. 1971. Balanced male steriles and dominant pre-flowering selective genes for use in hybrid seed production. p. 292-297. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
2. Eslick, R.F., and E.A. Hockett. 1972. Recombination values of four genes on chromosome 1. BGN 2:123-126.
3. Hockett, E.A. 1975. The genetic male sterile collection. BGN 5:84-86.
4. Hockett, E.A. 1979. The genetic male sterile collection. BGN 9:124-128.
5. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
6. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
7. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
8. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.

E.A. Hockett. 1971. BGN 1:184.

J.D. Franckowiak. 1997. BGN 26:317.

Stock number: BGS 370
Locus name: Male sterile genetic 15
Locus symbol: msg15

Previous nomenclature and gene symbolization:

Male sterile 15 = ms15 (1).

Monofactorial recessive (1).
Location is unknown.

Selfing - early tillers have 7.7% and late tillers have 46.2% at Bozeman, Montana, USA (1); only fully fertile plants were recovered from the attempt to develop a Bowman backcross-derived line.
Outcrossing - complete female fertility (1).
Stamens - anthers smaller than fertile sibs (1).
The phenotype of msg15.cn plants differ from other male sterile genetic mutants because late tillers are fertile (1). Anthesis is delayed in relationship to spike emergence from the flag leaf.

A spontaneous mutant in a selection from the cross Atlas/2*Kindred (CIho 13446) (1).

msg15.cn in Atlas/2*Kindred (MSS064) (1, 2).

msg15.cn in Atlas/2*Kindred (GSHO 1815).

1. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
2. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.

E.A. Hockett. 1971. BGN 1:184-185.

J.D. Franckowiak. 1997. BGN 26:318.

Stock number: BGS 371
Locus name: Male sterile genetic 16
Locus symbol: msg16

Previous nomenclature and gene symbolization:

Male sterile 16 = ms16 (6).

Monofactorial recessive (5).
Located in chromosome 5HS [7S] (2, 6), 8.0 to 15.0 cM distal from the breakpoint in translocation stock T2-7a (9, 10).

Selfing - none (3, 5).
Outcrossing - complete female fertility (5).
Stamens - anthers are almost equal in size to fertile sibs, stomium present, and filament elongation (5, 11).
Pollen - partial staining (15 to 35%) from both field and greenhouse grown plants, normal-appearing grains occur in some samples (11). Pollen appears immature based on staining with Fast Blue B, grains lack a pore or aperture, the exine is thinner, not distinctly two-layered, and bears fewer spicules than normal pollen (1). Tapetal tissue degeneration was observed at the free microspore stage (8).

A spontaneous mutant in Betzes (PI 129430) (5).

msg16.co in Betzes (MSS065) (5, 7); msg16.bi in Betzes (MSS323) (3, 4, 7).

msg16.co in Betzes (GSHO 1816); msg16.co in Bowman (PI 483237)*7 (GSHO 2116).

1. Ahokas, H. 1976. Male sterile mutants of barley. III. Additional inaperturate mutants. BGN 6:4-6.
2. Hayes, J.D., and M.S. Rana. 1966. Investigations on genetic resistance to chemicals in spring barley. p. 47-48. Welsh Plant Breed. Station, Aberystwyth, Report for 1965.
3. Hockett, E.A. 1972. Coordinator's report on the genetic male sterile barley collection. BGN 2:139-144.
4. Hockett, E.A. 1975. The genetic male sterile collection. BGN 5:84-86.
5. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
6. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
7. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
8. Mian, H.R., J. Kuspira, G.W.R. Walker, and N. Muntjewerff. 1981. Macromolecular changes and submicroscopic structure in the differentiation of msg16 male-sterile barley anthers. p. 804-813. In M.J.C. Asher, R.P. Ellis, A.M. Hayter, and R.N.H. Whitehouse (eds.) Barley Genetics IV. Proc. Fourth Int. Barley Genet. Symp., Edinburgh. Edinburgh Univ. Press, Edinburgh.
9. Ramage, R.T., and R.F. Eslick. 1975. Translocation linkage tests -- T2-7a X male sterile genes. BGN 5:46-48.
10. Ramage, R.T., M. Paluska, and G.A. Wiebe. 1973. Genetics and cytology of the translocation T2-7a. BGN 3:47-49.
11. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.

E.A. Hockett. 1971. BGN 1:185.

J.D. Franckowiak and U. Lundqvist. 1997. BGN 26:319.

Stock number: BGS 372
Locus name: Male sterile genetic 17
Locus symbol: msg17

Previous nomenclature and gene symbolization:

Male sterile 17 = ms17 (1).

Monofactorial recessive (1).
Location is unknown.

Selfing - 1% at Bozeman, Montana and 3% at Tucson, Arizona, USA (1).
Outcrossing - complete female fertility (1).
Stamens - anthers smaller than fertile sib, no stomium or filament elongation (3).
Pollen - no staining with 2,3,5-triphenyltetrazolium, but staining with acetocarmine, some normal-appearing grains (3).

A spontaneous mutant in Compana (PI 539111) (1).

msg17.cp in Compana (MSS066) (1, 2).

msg17.cp in Compana (GSHO 1817); msg17.cp in Bowman (PI 483237)*7 (GSHO 2301).

1. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
2. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
3. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.

E.A. Hockett. 1971. BGN 1:186.

J.D. Franckowiak. 1997. BGN 26:320.

Stock number: BGS 373
Locus name: Male sterile genetic 18
Locus symbol: msg18

Previous nomenclature and gene symbolization:

Male sterile 18 = ms18 (4).

Monofactorial recessive (4).
Located in chromosome 5HL [7L] (1), about 37.0 cM distal from the centromere (6).

Selfing - none (4).
Outcrossing - complete female fertility (4).
Stamens - anthers smaller than fertile sib, no stomium or filament elongation (7).
Pollen - no staining with 2,3,5-triphenyltetrazolium, but staining with acetocarmine, some normal-appearing grains (7).

A spontaneous mutant in Compana (PI 539111) (4).

msg18.cq in Compana (MSS067) (4, 5); msg18.z (MSS093) in Betzes (PI 129430) (3, 5); msg18.am (MSS304) in Betzes (2, 5).

msg18.cq in Compana (GSHO 1818); msg18.cq in Bowman (PI 483237)*7 (GSHO 2117).

1. Eslick, R.F. 1971. Balanced male steriles and dominant pre-flowering selective genes for use in hybrid seed production. p. 292-297. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
2. Hockett, E.A. 1976. The genetic male sterile collection. BGN 6:108.
3. Hockett, E.A. 1977. The genetic male sterile collection. BGN 7:97-100.
4. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
5. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
6. Ramage, R.T., and R.F. Eslick. 1975. Translocation linkage tests -- T2-7a X male sterile genes. BGN 5:46-48.
7. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.

E.A. Hockett. 1971. BGN 1:186-187.

J.D. Franckowiak. 1997. BGN 26:321.

Stock number: BGS 374
Locus name: Male sterile genetic 19
Locus symbol: msg19

Previous nomenclature and gene symbolization:

Male sterile 19 = ms19 (2).

Monofactorial recessive (2).
Located in chromosome 5HS [7S] (1, 3), between the centromere and the breakpoint in translocation stock T2-7a (5).

Selfing - none (2).
Outcrossing - complete female fertility (2).
Stamens - anthers smaller than fertile sib, no stomium or filament elongation (6).
Pollen - no staining with 2,3,5-triphenyltetrazolium, but staining with acetocarmine, some normal-appearing grains (6).

A spontaneous mutant in an introduction from Russia (CIho 14393) (2).

msg19.cr in an introduction from Russia (MSS068) (2, 4).

msg19.cr in an introduction from Russia (GSHO 1819); msg19.cr in Bowman (PI 483237)*6 (GSHO 2118).

1. Eslick, R.F. 1971. Balanced male steriles and dominant pre-flowering selective genes for use in hybrid seed production. p. 292-297. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
2. Hockett, E.A., and R.F. Eslick. 1968. Genetic male sterility in barley. I. Nonallelic genes. Crop Sci. 8:218-220.
3. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
4. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
5. Ramage, R.T., M. Paluska, and G.A. Wiebe. 1973. Genetics and cytology of the translocation T2-7a. BGN 3:47-49.
6. Roath, W.W., and E.A. Hockett. 1971. Genetic male sterility in barley. III. Pollen and anther characteristics. Crop Sci. 11:200-203.

E.A. Hockett. 1971. BGN 1:187.

J.D. Franckowiak. 1997. BGN 26:322.

Stock number: BGS 375
Locus name: Male sterile genetic 20
Locus symbol: msg20

Previous nomenclature and gene symbolization:

Male sterile ad = msg,,ad (1, 2, 4).
Male sterile 20 = ms 20 (1).

Monofactorial recessive (1).
Located in chromosome 1H [5] (1).

Selfing - none (1).
Outcrossing - complete female fertility (1).
Stamens - anthers smaller than fertile sib (1). Selfed seed set in the Bowman backcross-derived line is 50 to 75%, but both the original stock and the derived line have short awns (3/4 normal length).

An X-ray induced mutant in Hannchen (CIho 531) (1).

msg20.ad in Hannchen (MSS096) (1, 3).

msg20.ad in Hannchen (GSHO 2372); msg20.ad in Bowman (PI 483237)*5 (GSHO 2059).

1. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
2. Hockett, E.A., R.F. Eslick, D.A. Reid, and G.A. Wiebe. 1968. Genetic male sterility in barley. II. Available spring and winter stocks. Crop Sci. 8:754-755.
3. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
4. Robertson, D.W. 1971. Recent information of linkage and chromosome mapping. p. 220-242. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press.

E.A. Hockett. 1971. BGN 1:188.

T. Tsuchiya. 1982. BGN 12:107.
J.D. Franckowiak. 1997. BGN 26:323.

Stock number: BGS 376
Locus name: Male sterile genetic 21
Locus symbol: msg21

Previous nomenclature and gene symbolization:

Male sterile q = msg,,q (1, 2, 4).
Male sterile 21 = ms 21 (1).

Monofactorial recessive (1).
Location is unknown.

Selfing - 3.8% at Bozeman, Montana and 2.9% at Tucson, Arizona, USA (1, 2).
Outcrossing - complete female fertility (1).
Stamens - anthers almost equal in size to fertile sibs (1). The presumed Bowman backcross-derived line has 70 to 85% selfed seed set.

A spontaneous mutant in a Midwest Bulk (CIho 13640) (2).

msg21.q in CIho 13640 (MSS085) (1, 3).

msg21.q in CIho 13640 (GSHO 2373); msg21.q in Bowman (PI 483237)*7 (GSHO 2302).

1. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
2. Hockett, E.A., R.F. Eslick, D.A. Reid, and G.A. Wiebe. 1968. Genetic male sterility in barley. II. Available spring and winter stocks. Crop Sci. 8:754-755.
3. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
4. Robertson, D.W. 1971. Recent information of linkage and chromosome mapping. p. 220-242. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.

E.A. Hockett. 1971. BGN 1:188-189.

T. Tsuchiya. 1982. BGN 12:108.
J.D. Franckowiak. 1997. BGN 26:324.

Stock number: BGS 377
Locus name: Shrunken endosperm 1
Locus symbol: seg1

Previous nomenclature and gene symbolization:

Shrunken endosperm = se1 (2).

Monofactorial recessive (1).
Located in chromosome 7HL [1L] (1), linked to the msg23 (male sterile genetic 23) locus (3).

Seed is long and thin and the 100-kernel weight is about 33% of normal. Good stands can be established in the field if optimum environmental conditions prevail during germination and emergence (1, 3). This mutant is associated with an increase in percentage lysine in the protein (3).

A spontaneous mutant in Betzes (PI 129430) (1).

seg1.a in Betzes (1, 2).

seg1.a in Betzes (GSHO 750); seg1.a in Bowman (PI 483237)*7 (GSHO 1852).

1. Jarvi, A.J. 1970. Shrunken endosperm mutants in barley, Hordeum vulgare. Ph.D. Thesis. Montana State Univ., Bozeman.
2. Jarvi, A.J., and R.F. Eslick. 1971. BGS 377, Normal vs. shrunken endosperm, se1. BGN 1:190.
3. Jarvi, A.J., and R.F. Eslick. 1975. Shrunken endosperm mutants in barley. Crop Sci. 15:363-366.

A.J. Jarvi and R.F. Eslick. 1971. BGN 1:190.

R.F. Eslick. 1976. BGN 6:135.
T. Tsuchiya. 1980. BGN 10:124.
J.D. Franckowiak. 1997. BGN 26:325.

Stock number: BGS 378
Locus name: Shrunken endosperm 2
Locus symbol: seg2

Previous nomenclature and gene symbolization:

Shrunken endosperm = se2 (2).

Monofactorial recessive (1).
Located in chromosome 7HS [1S] (1, 5), about 5.7 cM proximal from the msg10 (male sterile genetic 10) locus (3), and about 2.8 cM proximal from the sex6 (shrunken endosperm xenia 6) locus (4).

Seed size is greatly reduced with a 100-kernel weight about 15% of normal. Fertility is normal. Little or no starch is deposited in the seed that develops on homozygous plants. When seed is germinated on blotters in a 10% sucrose solution, seedling establishment was about 25% after transplanting to soil (1, 3). The line maintained as the seg2.b stock (GSHO 751) has nearly normal appearing endosperm.

A spontaneous mutant in Betzes (PI 129430) (1).

seg2.b in Betzes (1, 2).

seg2.b from Betzes in six-rowed line (GSHO 751); seg2.b in a balanced tertiary trisomic (BBT) stock T16c (5).

1. Jarvi, A.J. 1970. Shrunken endosperm mutants in barley, Hordeum vulgare. Ph.D. Thesis. Montana State Univ., Bozeman.
2. Jarvi, A.J., and R.F. Eslick. 1971. BGS 378, Normal vs. shrunken endosperm, se2. BGN 1:190-191.
3. Jarvi, A.J., and R.F. Eslick. 1975. Shrunken endosperm mutants in barley. Crop Sci. 15:363-366.
4. Netsvetaev, V.P. 1990. [Location of a shrunken endosperm gene, sex6, in barley.] Nauchno-Tekhn. Byull' VSGI, Odessa. No. 1 (75):31-35. [In Russian.]
5. Ramage, R.T. 1971. Translocations and balanced tertiary trisomics. BGN 1:74-80.

A.J. Jarvi and R.F. Eslick. 1971. BGN 1:190-191.

R.F. Eslick. 1976. BGN 6:136.
T. Tsuchiya. 1980. BGN 10:125.
J.D. Franckowiak. 1997. BGN 26:326.

Stock number: BGS 379
Locus name: Shrunken endosperm 3
Locus symbol: seg3

Previous nomenclature and gene symbolization:

Shrunken endosperm = se3 (2).

Monofactorial recessive (1).
Located in chromosome 3H, over 30.8 cM from the centromere (3).

Seed size is reduced to about 33% of normal when grown under field conditions. Seeds are long and thin similar to those from seg1 plants; seeds are viable and good stand establishment is possible (3).

A spontaneous mutant in Compana (PI 539111) (1).

seg3.c in Compana (1, 2).

seg3.c in Compana (GSHO 752); seg3.c in Bowman (PI 483237)*7 (GSHO 1957).

1. Jarvi, A.J. 1970. Shrunken endosperm mutants in barley, Hordeum vulgare. Ph.D. Thesis. Montana State Univ., Bozeman.
2. Jarvi, A.J., and R.F. Eslick. 1971. BGS 379, Normal vs. shrunken endosperm, se3. BGN 1:191.
3. Jarvi, A.J., and R.F. Eslick. 1975. Shrunken endosperm mutants in barley. Crop Sci. 15:363-366.

A.J. Jarvi and R.F. Eslick. 1971. BGN 1:191.

R.F. Eslick. 1976. BGN 6:137.
T. Tsuchiya. 1980. BGN 10:126.
J.D. Franckowiak. 1997. BGN 26:327.

Stock number: BGS 380
Locus name: Shrunken endosperm 4
Locus symbol: seg4

Previous nomenclature and gene symbolization:

Shrunken endosperm = se4 (2).

Monofactorial recessive (1).
Located in chromosome 7HL [1L] (1), over 34.0 cM from the centromere (3).

Under field conditions in the original mutant, seed size is reduced to about 38% of normal and seed set is about 50% of normal. Stand establishment is poor under field conditions (1, 3). Xenia effects were not observed in the original study (1), but the stock presently held as GSHO 753 and the Bowman backcross-derived line show xenia effects in hybrid plants. The lemma side of the kernel has a very distinct depression in mature seed.

A spontaneous mutant in Compana (PI 539111) (1).

seg4.d in Compana (1, 2).

seg4.d in Compana (GSHO 753); seg4.d in Bowman (PI 483237)*5 (GSHO 1853).

1. Jarvi, A.J. 1970. Shrunken endosperm mutants in barley, Hordeum vulgare. Ph.D. Thesis. Montana State Univ., Bozeman.
2. Jarvi, A.J., and R.F. Eslick. 1971. BGS 380, Normal vs. shrunken endosperm, se4. BGN 1:192.
3. Jarvi, A.J., and R.F. Eslick. 1975. Shrunken endosperm mutants in barley. Crop Sci. 15:363-366.

A.J. Jarvi and R.F. Eslick. 1971. BGN 1:192.

R.F. Eslick. 1976. BGN 6:138.
T. Tsuchiya. 1980. BGN 10:127.
J.D. Franckowiak. 1997. BGN 26:328.

Stock number: BGS 381
Locus name: Shrunken endosperm 5
Locus symbol: seg5

Previous nomenclature and gene symbolization:

Shrunken endosperm = se5 (2).

Monofactorial recessive (1).
Located in chromosome 7HS [1S] (1), probably in the centromeric region of the short arm (3).

Seed size is reduced in homozygotes with the 100-kernel weight being about 50% of normal. However, seed size is quite variable within a spike. Seed set is reduced to about 20%, probably due to seed abortion. The caryopsis frequently exceeds the lemma and palea in length. The combination of caryopsis length and reduced seed set is used to classify mutant plants (1, 3).

A spontaneous mutant in a Sermo/7*Glacier (CIho 6976) line (1).

seg5.e in a Sermo/7*Glacier line (1, 2).

seg5.e in a Sermo/7*Glacier line (GSHO 754).

1. Jarvi, A.J. 1970. Shrunken endosperm mutants in barley, Hordeum vulgare. Ph.D. Thesis. Montana State Univ., Bozeman.
2. Jarvi, A.J., and R.F. Eslick. 1971. BGS 381, Normal vs. shrunken endosperm, se5. BGN 1:192-193.
3. Jarvi, A.J., and R.F. Eslick. 1975. Shrunken endosperm mutants in barley. Crop Sci. 15:363-366.

A.J. Jarvi and R.F. Eslick. 1971. BGN 1:192-193.

R.F. Eslick. 1976. BGN 6:139.
T. Tsuchiya. 1980. BGN 10:128.
J.D. Franckowiak. 1997. BGN 26:329.

Stock number: BGS 382
Locus name: Shrunken endosperm xenia 1
Locus symbol: sex1

Previous nomenclature and gene symbolization:

Shrunken endosperm 6 = se6 (1, 5).
Shrunken endosperm 6 = se6f (5).
High lysine 5 = lys5 (7, 8).

Monofactorial recessive (1, 5, 7).
Located in chromosome 6HL (2, 7, 9), about 5.6 cM proximal from the msg36 (male sterile genetic 36) locus (1, 3).

After the hard dough stage, kernels develop a depression in the center of the lemma which becomes progressively more distinct with maturity. The 100-kernel weight of the mutant is about 75% of normal. The mature endosperm appears harder than normal endosperm. The mutant expresses xenia and kernels from heterozygous plants can be classified as normal or shrunken with an expected 3:1 ratio. Establishment and growth of mutant plants is normal (4, 6). Mutant plants have a higher lysine content in the endosperm than normal sibs (8, 11).

A spontaneous mutant in Compana (PI 539111) (4, 5).

sex1.a in Compana (4, 9); lys5.f (Risø 13) in Bomi (PI 43371), sex1.d or lys5.h (Risø 86), sex1.e or lys5.g (Risø 29, GSHO 2474) in Carlsberg II (CIho 10114) (8, 9).

sex1.a in Compana (GSHO 755); sex1.e in Bowman (PI 483237)*3 (GSHO 2071).

1. Eslick, R.F., and M.N. Ries. 1976. Positioning sex1 on chromosome 6. BGN 6:21-22.
2. Falk, D.E. 1994. Creation of a marked telo 6S trisomic for chromosome 6. BGN 23:32.
3. Falk, D.E., M.J. Swartz, and K.J. Kasha. 1980. Linkage data with genes near the centromere of barley chromosome 6. BGN 10:13-16.
4. Jarvi, A.J. 1970. Shrunken endosperm mutants in barley, Hordeum vulgare. Ph.D. Thesis. Montana State Univ., Bozeman.
5. Jarvi, A.J., and R.F. Eslick. 1971. BGS 382, Normal vs. shrunken endosperm, se6. BGN 1:193.
6. Jarvi, A.J., and R.F. Eslick. 1975. Shrunken endosperm mutants in barley. Crop Sci. 15:363-366.
7. Jensen, J. 1979. Chromosomal location of one dominant and four recessive high-lysine genes in barley mutants. Vol. 1. p. 89-96. In Seed Protein Improvement in Cereals and Grain Legumes. Proc. Int. Symp., Neuherburg, Germany, 1978. Int. Atomic Energy Agency, Vienna.
8. Jensen, J., and H. Doll. 1979. Gene symbols for barley high-lysine mutants. BGN 9:33-37.
9. Ullrich, S.E., and R.F. Eslick. 1978. Allelism evidence for high lysine, shrunken endosperm xenia (sex) mutants. BGN 8:109-112.
10. Ullrich, S.E., and R.F. Eslick. 1978. Chromosome location evidence for Risø induced high lysine shrunken endosperm mutants of barley. BGN 8:114-125.
11. Ullrich, S.E., and R.F. Eslick. 1978. Lysine and protein characterization of spontaneous shrunken endosperm mutants in barley. Crop Sci. 18:809-812.

A.J. Jarvi and R.F. Eslick. 1971. BGN 1:193.

R.F. Eslick and E.A. Hockett. 1976. BGN 6:140.
T. Tsuchiya. 1980. BGN 10:129.
J.D. Franckowiak. 1997. BGN 26:330.

Stock number: BGS 383
Locus name: Male sterile genetic 22
Locus symbol: msg22

Previous nomenclature and gene symbolization:

Male sterile e = msg,,e (4, 5).
Male sterile 22 = ms22 (1, 4).

Monofactorial recessive (4).
Located in chromosome 7H [1] (4).

Selfing - none (4).
Outcrossing - complete female fertility (4).
Stamens - anthers smaller than fertile sibs, no stomium or filament elongation (2).

A spontaneous mutant in a selection from Glacier/Compana (CIho 10861) (5).

msg22.e in a selection from Glacier/Compana (MSS073) (5, 6); msg22.fc (MSS404) in Proctor (PI 280420), msg22.fo (MSS416) in Zephyr (PI 339815) (2, 3).

msg22.e in a selection from Glacier/Compana (GSHO 741, GSHO 2374); msg22.e in Bowman (PI 483237)*7 (GSHO 1857).

1. Hockett, E.A. 1972. Coordinator's report on the genetic male sterile barley collection. BGN 2:139-144.
2. Hockett, E.A. 1984. Coordinator's report. The genetic male sterile barley collection. BGN 14:70-75.
3. Hockett, E.A. 1991. The identification of eight new loci and allelism of 14 additional mutants. BGN 20:37-40.
4. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
5. Hockett, E.A., R.F. Eslick, D.A. Reid, and G.A. Wiebe. 1968. Genetic male sterility in barley. II. Available spring and winter stocks. Crop Sci. 8:754-755.
6. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.

E.A. Hockett. 1972. BGN 2:178 as BGS 377, Male sterile 22, ms22.
E.A. Hockett. 1973. BGN 3:121.

J.D. Franckowiak. 1997. BGN 26:331.

Stock number: BGS 384
Locus name: Male sterile genetic 23
Locus symbol: msg23

Previous nomenclature and gene symbolization:

Male sterile b = msg,,b (3, 4).
Male sterile 23 = ms23 (1, 3).

Monofactorial recessive (3).
Located in chromosome 7HL [1L], not near the centromere (3).

Selfing - none in msg23.b and msg23.bg (3), but 1.9 to 13.6% in msg23.y (2).
Outcrossing - complete female fertility (3).
Stamens - anthers smaller than fertile sibs, no stomium or filament elongation (3).

A spontaneous mutant in Betzes (PI 129430) (4).

msg23.b in Betzes (MSS071) (4, 5); msg23.y in Betzes (MSS092) (2, 3, 5); msg23.bg in Betzes (MSS321) (1, 2, 5).

msg23.b in Betzes (GSHO 2375); msg23.b in Bowman (PI 483237)*8 (GSHO 1867).

1. Hockett, E.A. 1972. Coordinator's report on the genetic male sterile barley collection. BGN 2:139-144.
2. Hockett, E.A. 1979. The genetic male sterile collection. BGN 9:124-128.
3. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
4. Hockett, E.A., R.F. Eslick, D.A. Reid, and G.A. Wiebe. 1968. Genetic male sterility in barley. II. Available spring and winter stocks. Crop Sci. 8:754-755.
5. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.

E.A. Hockett. 1972. BGN 2:179 as BGS 378, Male sterile 23, ms23.
E.A. Hockett. 1973. BGN 3:122.

J.D. Franckowiak. 1997. BGN 26:332.

Stock number: BGS 385
Locus name: Male sterile genetic 24
Locus symbol: msg24

Previous nomenclature and gene symbolization:

Male sterile v = msg,,v (5, 6).
Male sterile 24 = ms24 (2, 5).

Monofactorial recessive (5).
Located in chromosome 4HL (5), over 11.0 cM proximal from the Blx1 (non-blue aleurone xenia 1) locus (8).

Selfing - none (2, 5).
Outcrossing - complete female fertility (5).
Stamens - anthers smaller than fertile sibs, no stomium or filament elongation (3).

A spontaneous mutant in Betzes (PI 129430) (6).

msg24.j (MSS078) in Betzes/Domen (CIho 13639) (6); msg24.v in Betzes (MSS089) (6, 7); msg24.ak in Betzes (MSS302) (2, 6); msg24.an in Betzes (MSS305) (2, 7); msg24.at (MSS312) in OAC21 (CIho 1470), msg24.bc in Betzes (MSS317) (3, 7); msg24.hg in an unknown cultivar (MSS460) (1, 4).

msg24.v in Betzes (GSHO 2376); msg24.v in Bowman (PI 483237)*7 (GSHO 2018).

1. Dawi, D.A., and C.A. Foster. 1983. Allelism studies of new genetic male sterile barley stocks in the WPBS collection. BGN 13:9-11.
2. Hockett, E.A. 1972. Coordinator's report on the genetic male sterile barley collection. BGN 2:139-144.
3. Hockett, E.A. 1975. The genetic male sterile collection. BGN 5:84-86.
4. Hockett, E.A. 1984. Coordinator's report. The genetic male sterile barley collection. BGN 14:70-75.
5. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
6. Hockett, E.A., R.F. Eslick, D.A. Reid, and G.A. Wiebe. 1968. Genetic male sterility in barley. II. Available spring and winter stocks. Crop Sci. 8:754-755.
7. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.
8. Kushnak, G.D. 1972. Linkage of Bl and ms-v on chromosome 4. BGN 2:45-46.

E.A. Hockett. 1973. BGN 3:123.

J.D. Franckowiak. 1997. BGN 26:333.

Stock number: BGS 386
Locus name: Desynapsis 3
Locus symbol: des3

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2, 3).
Location is unknown.

The chromosomes are paired during the pachytene stage of microsporogenesis, but they undergo desynapsis during diplotene and diakinesis. The degree of desynapsis is 0.9 +/- 1.7, ranging from 7 ring bivalents (d = 0) to 4 ring bivalents plus 2 rod bivalents and 2 univalents (d = 4). Ovule fertility ranges from 1% to 33%, much too low to be explained by the low degree of desynapsis observed in microsporocytes (1).

A spontaneous mutant in Betzes (PI 129430) (2, 3).

des3.c in Betzes (2, 3).

des3.c in Betzes (GSHO 594).

1. Hernandez-Soriano, J.M. 1973. Desynaptic mutants in Betzes barley. M.S. Thesis. Univ. of Arizona, Tucson.
2. Ramage, R.T., and J.M. Hernandez-Soriano. 1971. Desynaptic genes in Betzes barley. BGN 1:38.
3. Ramage, R.T., and J.M. Hernandez-Soriano. 1972. Desynaptic genes in barley. BGN 2:65-68.

J.M. Hernandez-Soriano, R.T. Ramage, and R.F. Eslick. 1973. BGN 3:126.

J.D. Franckowiak. 1997. BGN 26:334.

Stock number: BGS 387
Locus name: Desynapsis 8
Locus symbol: des8

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (2, 3).
Location is unknown.

The chromosomes are paired during pachytene and undergo desynapsis during diplotene. The degree of desynapsis is 7.9 +/- 2.3, ranging from 7 ring bivalents (d = 0) to 14 univalents (d = 14). Many univalents split longitudinally during anaphase I. Lagging chromosomes and micronuclei are frequently observed at telophase I. Microspore quartets contain an average of 1.2 micronuclei per quartet with a range of 0 to 10. Ovule fertility is about 43% in the des8.k stock, but about 1% in the des8.l stock (1).

A spontaneous mutant in Betzes (PI 129430) (2, 3).

des8.k, des8.l in Betzes (2, 3).

des8.k in Betzes (GSHO 599).

1. Hernandez-Soriano, J.M. 1973. Desynaptic mutants in Betzes barley. M.S. Thesis. Univ. of Arizona, Tucson.
2. Ramage, R.T., and J.M. Hernandez-Soriano. 1971. Desynaptic genes in Betzes barley. BGN 1:38.
3. Ramage, R.T., and J.M. Hernandez-Soriano. 1972. Desynaptic genes in barley. BGN 2:65-68.

J.M. Hernandez-Soriano, R.T. Ramage, and R.F. Eslick. 1973. BGN 3:131.

J.D. Franckowiak. 1997. BGN 26:335.

Stock number: BGS 388
Locus name: Desynapsis 9
Locus symbol: des9

Previous nomenclature and gene symbolization:

Desynapsis n = des,,n (1, 3).

Monofactorial recessive (1, 3).
Location is unknown.

The chromosomes are paired during pachytene and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 0.9, ranging from 7 ring bivalents (d = 0) to 5 rod bivalents (d = 5). Ovule fertility is about 90% (2).

A spontaneous mutant in Betzes (PI 129430) (3).

des9.n in Betzes (1, 3).

des9.n in Betzes (GSHO 600).

1. Hernandez-Soriano, J.M., and R.T. Ramage. 1973. Coordinator's report: Desynaptic genes. BGN 3:91.
2. Hernandez-Soriano, J.M., and R.T. Ramage. 1974. BGS 388, Normal vs. desynapsis 9, des9. BGN 4:137.
3. Ramage, R.T., and J.M. Hernandez-Soriano. 1972. Desynaptic genes in barley. BGN 2:65-68.

J.M. Hernandez-Soriano and R.T. Ramage. 1974. BGN 4:137.

J.D. Franckowiak. 1997. BGN 26:336.

Stock number: BGS 389
Locus name: Desynapsis 10
Locus symbol: des10

Previous nomenclature and gene symbolization:

Desynapsis p = des,,p (1, 3).

Monofactorial recessive (1, 3).
Location is unknown.

The chromosomes are paired during pachytene and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 2.0, ranging from 7 ring bivalents (d = 0) to 3 ring bivalents plus 2 rod bivalents plus 4 univalents (d = 6). Ovule fertility is 60 to 80% (2).

A spontaneous mutant in Betzes (PI 129430) (3).

des10.p in Betzes (1, 3).

des10.p in Betzes (GSHO 601).

1. Hernandez-Soriano, J.M., and R.T. Ramage. 1973. Coordinator's report: Desynaptic genes. BGN 3:91.
2. Hernandez-Soriano, J.M., and R.T. Ramage. 1974. BGS 389, Normal vs. desynapsis 10, des10. BGN 4:138.
2. Ramage, R.T., and J.M. Hernandez-Soriano. 1972. Desynaptic genes in barley. BGN 2:65-68.

J.M. Hernandez-Soriano and R.T. Ramage. 1974. BGN 4:138.

J.D. Franckowiak. 1997. BGN 26:337.

Stock number: BGS 390
Locus name: Desynapsis 11
Locus symbol: des11

Previous nomenclature and gene symbolization:

Desynapsis r = des,,r (1, 3).

Monofactorial recessive (1, 3).
Location is unknown.

The chromosomes are paired during pachytene and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 5.7, ranging from 6 ring bivalents plus one rod bivalent (d = 1) to 2 rod bivalents plus 10 univalents (d = 12). Ovule fertility is about 40% (2). When crossed with non-allelic desynaptic mutants des4.d and des5.e, the F₁'s frequently show a low degree of desynapsis (up to 3 rod bivalents). When crossed with des7.j, the F₁'s show a low to medium degree of desynapsis (d = 1.8), ranging from 7 ring bivalents (d = 0) to 4 ring bivalents plus 4 rod bivalents (d = 5), and there are lagging chromosomes and micronuclei at telophase I (2).

A spontaneous mutant in Betzes (PI 129430) (3).

des11.r in Betzes (1, 3).

des11.r in Betzes (GSHO 602).

1. Hernandez-Soriano, J.M., and R.T. Ramage. 1973. Coordinator's report: Desynaptic genes. BGN 3:91.
2. Hernandez-Soriano, J.M., and R.T. Ramage. 1974. BGS 390, Normal vs. desynapsis 11, des11. BGN 4:139.
3. Ramage, R.T., and J.M. Hernandez-Soriano. 1972. Desynaptic genes in barley. BGN 2:65-68.

J.M. Hernandez-Soriano and R.T. Ramage. 1974. BGN 4:139.

J.D. Franckowiak. 1997. BGN 26:338.

Stock number: BGS 391
Locus name: Desynapsis 12
Locus symbol: des12

Previous nomenclature and gene symbolization:

Desynapsis s = des,,s (1, 3).

Monofactorial recessive (1, 3).
Location is unknown.

The chromosomes are paired during pachytene and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 4.7, ranging from 7 ring bivalents (d = 0) to 1 rod bivalent plus 12 univalents (d = 13). Ovule fertility is about 35% (2).

A spontaneous mutant in Betzes (PI 129430) (3).

des12.s, des12.w in Betzes (1, 3).

des12.s in Betzes (GSHO 603).

1. Hernandez-Soriano, J.M., and R.T. Ramage. 1973. Coordinator's report: Desynaptic genes. BGN 3:91.
2. Hernandez-Soriano, J.M., and R.T. Ramage. 1974. BGS 391, Normal vs. desynapsis 12, des12. BGN 4:140.
3. Ramage, R.T., and J.M. Hernandez-Soriano. 1972. Desynaptic genes in barley. BGN 2:65-68.

J.M. Hernandez-Soriano and R.T. Ramage. 1974. BGN 4:140.

J.D. Franckowiak. 1997. BGN 26:339.

Stock number: BGS 392
Locus name: Desynapsis 13
Locus symbol: des13

Previous nomenclature and gene symbolization:

Desynapsis t = des,,t (1, 3).

Monofactorial recessive (1, 3).
Location is unknown.

There are two types of microspore mother cells in mutant plants. About 58% have normal early prophases and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 0.8, ranging from 7 ring bivalents (d = 0) to 4 ring bivalents plus 2 rod bivalents plus 2 univalents (d = 4). The other type of microspore mother cells have chromosomes that appear "sticky" at metaphase I. The stickiness is accompanied by very elongated ring bivalents and by extensive chromosome fragmentation. Ovule fertility is about 20 to 30% (2).

A spontaneous mutant in Betzes (PI 129430) (3).

des13.t in Betzes (1, 3).

des13.t in Betzes (GSHO 604).

1. Hernandez-Soriano, J.M., and R.T. Ramage. 1973. Coordinator's report: Desynaptic genes. BGN 3:91.
2. Hernandez-Soriano, J.M., and R.T. Ramage. 1974. BGS 392, Normal vs. desynapsis 13, des13. BGN 4:141.
3. Ramage, R.T., and J.M. Hernandez-Soriano. 1972. Desynaptic genes in barley. BGN 2:65-68.

J.M. Hernandez-Soriano and R.T. Ramage. 1974. BGN 4:141.

J.D. Franckowiak. 1997. BGN 26:340.

Stock number: BGS 393
Locus name: Desynapsis 14
Locus symbol: des14

Previous nomenclature and gene symbolization:

Desynapsis u = des,,u (1, 3).

Monofactorial recessive (1, 3).
Location is unknown.

The chromosomes are paired during pachytene and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 4.8, ranging from 7 ring bivalents (d = 0) to 1 rod bivalent plus 12 univalents (d = 13). Lagging chromosomes and micronuclei are observed frequently at telophase I. Ovule fertility is about 35% (2).

A spontaneous mutant in Betzes (PI 129430) (2).

des14.u in Betzes (1, 3).

des14.u in Betzes (GSHO 605).

1. Hernandez-Soriano, J.M., and R.T. Ramage. 1973. Coordinator's report: Desynaptic genes. BGN 3:91.
2. Hernandez-Soriano, J.M., and R.T. Ramage. 1974. BGS 393, Normal vs. desynapsis 14, des14. BGN 4:142.
3. Ramage, R.T., and J.M. Hernandez-Soriano. 1972. Desynaptic genes in barley. BGN 2:65-68.

J.M. Hernandez-Soriano and R.T. Ramage. 1974. BGN 4:142.

J.D. Franckowiak. 1997. BGN 26:341.

Stock number: BGS 394
Locus name: Desynapsis 15
Locus symbol: des15

Previous nomenclature and gene symbolization:

Desynapsis x = des,,x (1, 3).

Monofactorial recessive (1, 3).
Location is unknown.

The chromosomes are paired during pachytene and undergo desynapsis during diplotene and diakinesis. The average degree of desynapsis is 7.5, ranging from 5 ring bivalents plus 2 rod bivalents (d = 2) to 1 rod bivalent plus 12 univalents (d = 13). Ovule fertility is about 25% (2).

A spontaneous mutant in Ingrid (CIho 10083) (1).

des15.x, des15.y in Ingrid (1, 3).

des15.x in Ingrid (GSHO 606).

1. Hernandez-Soriano, J.M., and R.T. Ramage. 1973. Coordinator's report: Desynaptic genes. BGN 3:91.
2. Hernandez-Soriano, J.M., and R.T. Ramage. 1975. BGS 394, Normal vs. desynapsis 15, des15. BGN 5:113.
3. Scheuring, J.R., and R.T. Ramage. 1975. Reports from coordinators: Desynaptic genes. BGN 5:86.

J.M. Hernandez-Soriano and R.T. Ramage. 1975. BGN 5:113.

J.D. Franckowiak. 1997. BGN 26:342.

Stock number: BGS 395
Locus name: Male sterile genetic 26
Locus symbol: msg26

Previous nomenclature and gene symbolization:

Male sterile u = msg,,u (5).

Monofactorial recessive (3, 4).
Located in chromosome 7HS [1S] (1, 2), linked to the nud (naked caryopsis) locus (1), and linked to the ant1 (anthocyanin-less 1) locus based on linkage drag (2).

Selfing - none (4).
Outcrossing - complete female fertility (4).
Stamens - anthers smaller than fertile sibs, no stomium or filament elongation (4).

A spontaneous mutant in Unitan (CIho 10421) (5).

msg26.u in Unitan (MSS088) (3, 5, 6).

msg26.u in Unitan (GSHO 745, GSHO 2378); msg26.u in Bowman (PI 483237)*6 (GSHO 1858).

1. Franckowiak, J.D. 1988. Mapping four male sterile genes on chromosome 1. Barley Newsl. 31:111.
2. Franckowiak, J.D. 1995. Notes on linkage drag in Bowman backcross derived lines of spring barley. BGN 24:63-70.
3. Hockett, E.A. 1974. The genetic male sterile collection. BGN 4:121-123.
4. Hockett, E.A., and R.F. Eslick. 1971. Genetic male-sterile genes useful in hybrid barley production. p. 298-307. In R.A. Nilan (ed.) Barley Genetics II. Proc. Second Int. Barley Genet. Symp., Pullman, WA, 1969. Washington State Univ. Press, Pullman.
5. Hockett, E.A., R.F. Eslick, D.A. Reid, and G.A. Wiebe. 1968. Genetic male sterility in barley. II. Available spring and winter stocks. Crop Sci. 8:754-755.
6. Hockett, E.A., and D.A. Reid. 1981. Spring and winter genetic male-sterile barley stocks. Crop Sci. 21:655-659.

E.A. Hockett. 1974. BGN 4:136 as BGS 387, Male sterile genetic 26, msg26.
E.A. Hockett. 1975. BGN 5:170.

J.D. Franckowiak. 1997. BGN 26:343.

Stock number: BGS 396
Locus name: Shrunken endosperm 6
Locus symbol: seg6

Previous nomenclature and gene symbolization:

Shrunken endosperm = se6 (2).

Monofactorial recessive (1).
Located in chromosome 3HL (3).

Seed size is reduced, but the degree reduction is affected by environment. Seed weights of 25, 50, and 75% of normal are reported for plants grown in the field in Arizona, in the field in Montana, and in the greenhouse in Arizona, USA, respectively (3). Pollen mother cell meiosis and pollen fertility are normal. Seed from seg6.g plants can be used to establish stands under field conditions (3).

A spontaneous mutant in Ingrid (CIho 10083) (2).

seg6.f in an unknown hybrid, seg6.g in Ingrid (2).

seg6.g in Ingrid (GSHO 2467); seg6.g in Bowman (PI 483237)*3 (GSHO 1975).

1. Ramage, R.T. 1983. Chromosome location of shrunken endosperm mutants seg6g and seg8k. BGN 13:64-65.
2. Ramage, R.T., and R.F. Eslick. 1975. BGS 396, Shrunken endosperm, xenia, se6. BGN 5:114.
3. Ramage, R.T., and J.F. Scheuring. 1976. Shrunken endosperm mutants seg6 and seg7. BGN 6:59-60.

R.T. Ramage and R.F. Eslick. 1975. BGN 5:114.

R.T. Ramage and R.F. Eslick. 1976. BGN 6:141.
T. Tsuchiya. 1980. BGN 10:130.
R.T. Ramage. 1983. BGN 13:115.
J.D. Franckowiak. 1997. BGN 26:344.

Stock number: BGS 397
Locus name: Shrunken endosperm 7
Locus symbol: seg7

Previous nomenclature and gene symbolization:

Shrunken endosperm = se7 (2).

Monofactorial recessive (1).
Location is unknown.

Seed size is reduced less than other seg mutants, but the degree reduction is affected by environment. Seed weights of 40, 75, and 90% of normal are reported for plants grown in the field in Arizona, in the field in Montana, and in the greenhouse in Arizona, USA, respectively (3). Pollen mother cell meiosis and pollen fertility are normal. Seed from seg7.h plants can be used to establish stands under field conditions (3).

A spontaneous mutant in Ingrid (CIho 10083) (2).

seg7.h in Ingrid (2).

seg7.h in Ingrid (GSHO 2468); seg7.h in Bowman (PI 483237)*2 (GSHO 2352).

1. Ramage, R.T., and R.F. Eslick. 1975. BGS 397, Shrunken endosperm, xenia, se7. BGN 5:115.
2. Ramage, R.T., and J.F. Scheuring. 1976. Shrunken endosperm mutants seg6 and seg7. BGN 6:59-60.

R.T. Ramage and R.F. Eslick. 1975. BGN 5:115.

R.T. Ramage and R.F. Eslick. 1976. BGN 6:142.
T. Tsuchiya. 1980. BGN 10:131.
J.D. Franckowiak. 1997. BGN 26:345.

Stock number: BGS 399
Locus name: Eceriferum-d
Locus symbol: cer-d

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (1).
Location is unknown.

Surface wax coating on the spike is reduced (wax code + ++ ++) (1). A translocation is present in the mutant stock cer-d.5, but it is not linked to the cer-d locus. The original stock has a lax spike and heads early (1, 7). The Bowman backcross-derived line has narrow leaves and reduced vigor, and heads a few days earlier than Bowman.

An X-ray induced mutant in Bonus (PI 189763) (1).
Mutational events:
cer-d.5 (trans), -d.40, -d.104 in Bonus (1, 2); cer-d.111, -d.149 in Bonus (2, 7); cer-d.713 in Bonus (2); cer-d.809, -d.826 in Bonus (3); cer-d.965 in Bonus (4); cer-d.712, -d.1326 in Bonus (5); cer-d.1818, -d.1819, -d.1820 in Bonus (6).
Mutant used for description and seed stocks:
cer-d.5 in Bonus (GSHO 425); cer-d.5 in Bowman (PI 483237)*5 (GSHO 2189).

1. Lundqvist, U., and D. von Wettstein. 1962. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. Hereditas 48:342-362.
2. Lundqvist, U., and D. von Wettstein. 1971. Stock list for the eceriferum mutants. BGN 1:97-102.
3. Lundqvist, U., and D. von Wettstein. 1973. Stock list for the eceriferum mutants II. BGN 3:110-112.
4. Lundqvist, U., and D. von Wettstein. 1975. Stock list for the eceriferum mutants III. BGN 5:88-91.
5. Lundqvist, U., and D. von Wettstein. 1979. Stock list for the eceriferum mutants V. BGN 9:135-137.
6. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.
7. Lundqvist, U., P. von Wettstein-Knowles, and D. von Wettstein. 1968. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. II. Hereditas 59:473-504.

U. Lundqvist. 1975. BGN 5:117.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:346.

Stock number: BGS 400
Locus name: Eceriferum-e
Locus symbol: cer-e

Previous nomenclature and gene symbolization:

None.

Monofactorial recessive (3).
Located in chromosome 1HL [5L] (1, 12), close to the centromere and near the ert-b (erectoides-b) locus (2, 12, 13).

Surface wax coating on the spike appears absent or greatly reduced (wax code
-/+ ++ ++) (3). A translocation is present in the allele cer-e.18, but it is not linked to the cer-e locus (3). The Bowman backcross-derived line has less wax on the spike and upper leaf sheath (wax code - + ++).

An X-ray induced mutant in Bonus (PI 189763) (3).
Mutational events:
cer-e.8, -e.18 (trans), -e.34 in Bonus (3, 4); cer-e.90, -e.140 in Bonus, -e.272, -e.294, -e.388, -e.551, -e.560, -e.586, -e.607 in Foma (CIho 11333) (4); cer-e.273, -e.277, -e.296, -e.317, -e.321, -e.473, -e.480 in Foma (4, 11); cer-e.663, -e.692, -e.716, -e.717 in Bonus (4); cer-e.161 in Bonus, -e.577 in Foma, -e.790 in Bonus, -e.1089, -e.1106 in Kristina (5); cer-e.908, -e.962 in Bonus, -e.1162 in Kristina (6); cer-e.1335 in Bonus (7); cer-e.1366, -e.1404, -e.1425, -e.1426 in Bonus (8); cer-e.1403, -e.1709, -e.1715 in Bonus (9); cer-e.1519 in Nordal, -e.1760 in Bonus, -e.1873, -e.1878, -e.1880, -e.1884 in Sv 79353 (10).
Mutant used for description and seed stocks.
cer-e.8 in Bonus (GSHO 1518); cer-e.8 in Bowman (PI 483237)*7 (GSHO 2048).

1. Fester, T., and B. Søgaard. 1969. The localization of eceriferum loci in barley. Hereditas 61:327-337.
2. Jensen, J. 1974. Coordinator's report: Chromosome 5. BGN 4:102-106.
3. Lundqvist, U., and D. von Wettstein. 1962. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. Hereditas 48:342-362.
4. Lundqvist, U., and D. von Wettstein. 1971. Stock list for the eceriferum mutants. BGN 1:97-102.
5. Lundqvist, U., and D. von Wettstein. 1973. Stock list for the eceriferum mutants II. BGN 3:110-112.
6. Lundqvist, U., and D. von Wettstein. 1975. Stock list for the eceriferum mutants III. BGN 5:88-91.
7. Lundqvist, U., and D. von Wettstein. 1979. Stock list for the eceriferum mutants V. BGN 9:135-137.
8. Lundqvist, U., and D. von Wettstein. 1982. Stock list for the eceriferum mutants VI. BGN 12:169-172.
9. Lundqvist, U., and D. von Wettstein. 1985. Stock list for the eceriferum mutants VII. BGN 15:89-93.
10. Lundqvist, U., and D. von Wettstein. 1988. Stock list for the eceriferum mutants VIII. BGN 18:88-91.
11. Lundqvist, U., P. von Wettstein-Knowles, and D. von Wettstein. 1968. Induction of eceriferum mutants in barley by ionizing radiations and chemical mutagens. II. Hereditas 59:473-504.
12. Søgaard, B. 1971. Linkage studies on eceriferum mutants in barley. BGN 1:41-47.
13. Søgaard, B. 1973. Continued linkage studies on eceriferum mutants in barley. BGN 3:57-61.

U. Lundqvist. 1975. BGN 5:118.

U. Lundqvist and J.D. Franckowiak. 1997. BGN 26:347.