White Sands National Monument
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Checklist of Plants of the White Sands

A Discussion of Dunes Ecology With Revised Checklist

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INTRODUCTION

yucca.jpg-image of yucca in dunes

White Sands National Monument preserves a sea of graceful white gypsum sand dunes--a landscape of stark natural beauty. Life is difficult in the dune field, even for plants adapted to desert conditions. The dune field environment is unusually harsh: plants must endure burial by moving dunes, nutrient-poor gypsum soil, and extreme fluctuations of temperature. Only about 60 species of plants, one quarter of those growing in the adjacent Tularosa Basin, have found a way to survive in the dunes.

This guide combines previously compiled, revised checklists of plants of the White Sands. The following discussion of the ecological aspects of the dunes should help the observer to locate and identify individual species of plants. By placing the checklists in ecological context, it is hoped that the observer will come to think of each plant as a member of a life community interrelating with the dune movement.

ECOLOGICAL ASPECTS

Ecology is the study of the interrelationships between plants and animals and their physical surroundings. It is literally a study of the home or household of living things. In understanding the ecology of the White Sands, several rather distinct homes, or habitats, must be considered. Most of these habitats are named in terms of the physical surroundings because, as in most arid lands, the inorganic part of the environment is the most prominent and has the greatest influence on the organic parts.

Interdune flats: As the gypsum sand dunes move northwestward across the Tularosa Basin, pieces of the underlying desert floor are exposed between the individual dunes. Most of the plant species inside the dune field grow only in these interdune flats, where conditions are less hostile. Better soil and protection from wind and blowing sand allows plants to survive for a while-- until the next dune buries them.

The most showy wildflowers in the interdune flats include Centaury, a gentian with bright pink flowers; Sand Verbena, with its fragrant lilac-like smell; Stick-leaf, with yellow, star-like flowers; Woody Paperflower, which stands out against the white dunes in bright yellow clumps in the fall; and Yellow Evening Primrose.

Marginal Dunes: These extend into the dune field two or three miles from its southern and eastern boundary. Most of the dunes in this habitat are slow-moving, scattered and separated by large grassy, interdune areas. Although the dunes are still the prominent feature here, the effects of vegetation in slowing the rate of dune movement is very evident.

The marginal dunes themselves have become relatively heavily populated with flora able to withstand such physical conditions. There are eight species of plants that routinely grow on the marginal dunes. The Soaptree Yucca is found scattered throughout this part of the dune field. Yuccas that can be seen on the tops of dunes actually germinate in interdune areas. As a dune begins to bury them, the yuccas elongate their stems, growing upward as much as a foot per year, to keep their leaves above the sand.

Two large shrubs, Skunkbush Sumac and Hoary Rosemarymint, can also extend their stems and outgrow slow-moving marginal dunes. Their stems and roots can then anchor the dunes, further slowing dune movement and allowing other plants to take root on the relatively stable soil.

This developing plant community attracted animal life from the adjacent desert, which became fit to live on the dunes through evolutionary adaptation. Thus, the marginal dunes are now an ecological complex of unexpected variety and diversity.

Transverse and Barchan Dunes: In the center of the dune field, the physical forces of nature reign supreme. The paucity of plant life in the interior of the dune field is indicative of the harsh environmental conditions that prevail. Large transverse and barchan dunes creep forward many feet per year, overwhelming all plant life in their paths. Even the fast-growing yucca and rosemarymint cannot outgrow these dunes. No plants grow on the tops of the dunes, and only a few hardy species are able to live in the interdune flats until they are covered by sand. This interdune environment, known as the Abronia (Sand Verbena) association, is characterized by openings invaded first by Evening Primrose. These pioneers occupy the lee slope of the migrating dunes, the most recently created portion of the interdune flat. Moving out toward the center (older portion) of the flat, the Primrose is replaced by Indian Ricegrass and, later, Groundsel. The last plants to invade are Sand Verbena, Ephedra, Greenthread, and, finally, Alkali Sacaton.

Alkali Flat and Lake Lucero: The transverse-barchan dunes grade to the west into a narrow zone of embryonic dunes. The latter mark the eastern boundary of an ancient lake bed called the Alkali Flat. Here alkaline conditions prevent the growth of plant life except for a few scattered grasses and a scaly pseudo- evergreen known as Pickleweed. Lake Lucero, at the southern end of the Alkali Flat, occasionally contains standing water. There is little plant growth in the bed of Lake Lucero due to extreme alkaline conditions and infrequent flooding. However, alkaline- tolerant grasses sparsely fringe the shore of the lake.

Alluvial Fans: The alluvial fans at the base of the San Andres Mountains have coalesced to form a broad slope known as a bajada. The lowermost slopes of this extend into the monument and border the Alkali Flat on the west. The bajada is cut at frequent intervals by deep washes or arroyos that empty onto the Alkali Flat and Lake Lucero. Large Honey Mesquite hummocks are the dominant vegetative feature. The mesquite community soon gives way to Creosote Bush higher up on the slopes, near the monument boundary.

Saltbush Flats: The center of this high desert basin, the Tularosa, is vegetated mainly by Four Wing Saltbush and salt- tolerant bunch grasses. As grazing land, it is now very poor but, in the 1800's before the introduction of range cattle, it supposedly was predominately grassland having more of the appearance of plains than desert. A combination of drought and overgrazing may have allowed the hardy saltbush to take over. The eastern and southern edges of the marginal dunes are bordered by these saltbush flats. This grey-green monotony is broken by an occasional splash of bright red blossoms of the Claret Cup Hedgehog Cactus in the spring and stands of Golden Crownbeard in the fall.

Exotic plants: Tamarisk, or salt cedar, is a Mediterranean shrub introduced into North America. The aggressive tamarisk has spread throughout the southwest, growing thickly along streams, ponds and other seasonally-wet areas. At White Sands, tamarisk has invaded many interdune areas, where water is close to the surface. To protect native plants, the National Park Service is actively trying to control the spread of tamarisk within the monument. Other exotic plants now found in the park include Russian Thistle, or tumbleweed, and African Rue.

CLIMATIC FACTORS

This high desert basin, averaging 4,000 feet in elevation, is subject to harsh and sometimes rapidly changing climatic conditions. Summers are hot, averaging 95°F highs in July and August, with occasional readings over 100°F. Winter days are relatively mild, but nighttime temperatures routinely drop below freezing. Cold spells can send the mercury to below zero. The lowest temperature on record is a -25°F. Snowfall is infrequent, but heavy snows have occurred on occasion. Precipitation averages about eight inches per year, with most of this occurring during summer thunderstorms, often accompanied by hail.

Wind is the dominant climatic factor here, especially from February through May. The prevailing southwesterly winds blow unimpeded across the desert and at times reach gale proportions. Storms sometimes last for several days in the spring. This is the time of greatest dune movement, when living conditions for dune plant and animal communities become extremely harsh.

PLANT CHECKLIST

The following checklist has been compiled by various qualified individuals over the years since White Sands was established as a National Monument in 1933. Much of the work has been done by National Park Service personnel and volunteers. The latest revision was prepared by park volunteers Paul Shaw and Jeanine Derby. A Flora of New Mexico by W.C. Martin and C.R. Hutchins is the source for scientific names used in this checklist.

Separate checklists of birds and other animals of the white sands are available at the monument bookstore.


CHECKLIST FOR PLANTS

DIVISION - CYANOCHLORONTA (Cyanobacteria)

Oscillatoria sp.  

DIVISION - CHLOROPHYTA (Green Algae)

Chlamydomonas sp.  
Microcoleus paludosus  
Microcoleus vaginatus  
Nostoc spp.  
Palmogloea protuberans  
Pleconema nostocorum  
Schizothrix californica  
Schizothrix lamyi  
Scytonema hofmannii  

DIVISION - CHAROPHYTA (Stone Worts)

Nitella sp.  

DIVISION - EUGLENOPHYCOPHYTA (Euglendids)

Euglena sp.  

DIVISION - THALLOPHYTA (Lichens)

Dermatocarpon lachneum  
Fulgensia sp.  
Physia aipolia  
Psora sp.  
Psora decipiens  

DIVISION - SPERMATOPHYTA

GYMNOSPERMAE (Non-flowering Plants)

EPHEDRACEAE (Ephedra Family)

Ephedra torreyana

Mormon Tea or Jointfir

Ephedra trifurca Canatilla

ANGIOSPERMAE (Flowering Plants) MONOCOTYLEDONAE

CYPERACEAE (Sedge Family)

Scirpus paludosus Salt-Marsh Bulrush
Scirpus microcarpus Bulrush

JUNCACEAE (Rush Family)

Juncus balticus var. montanus Rush
Juncus mexicanus Rush

LILIACEAE (Lily Family)

Yucca elata

Soaptree Yucca

Yucca torreyi Torrey Yucca

POACEAE (Grass Family)

TRIBE (4) ABUNDINEAE

Arundo donax Giant Reed

TRIBE (8) STIPEAE

Oryzopsis hymenoides

Indian Ricegrass

TRIBE (13) ERAGROSTEAE

Muhlenbergia arenacea Ear Muhly
Muhlenbergia porteri Bush Muhly
Muhlenbergia pungens Sandhill Muhly
Scleropogon brevifolius Burrgrass
Sporobolus airoides

Alkali Sacaton

Sporobolus contractus Spike Dropseed
Sporobolus cryptandrus Sand Dropseed
Sporobolus flexuosis Mesa Dropseed
Sporobolus giganteus Giant Dropseed
Sporobolus nealleyi Gypgrass or Nealley Dropseed
Tridens pulchellus Fluff Grass

TRIBE (14) CHLORIDEAE

Bouteloua barbata Six-weeks Grama
Bouteloua breviseta Gyp Grama or Short- bristled Grama
Bouteloua gracilis Blue Grama
Chloris virgata Feather Fingergrass
Hilaria mutica Tobosa Grass

TRIBE (16) AELUROPODEAE

Distichlis stricta

Desert Saltgrass

TRIBE (19) ARISTIDEAE

Aristida adscensionis Six-weeks Three-awn

TRIBE (20) PANICEAE

Setaria macrcostachya Plains Bristlegrass

TRIBE (21) ANDROPOGONEAE

Andropogon scoparius

Little Bluestem

RUPPIACEAE (Ditch-grass Family)

Ruppia maritima Widgeongrass

TYPHACEAE (Cattail Family)

Typha angustifolia Narrow-leaved Cattail
Typha latifolia Common Cattail

ANGIOSPERMAE (Flowering Plants) DICOTYLEDONAE

AIZOACEAE (Carpet-weed Family)

Trianthema portulacastrum Horse Purslane

AMARANTHACEAE (Amaranth Family)

Amaranthus retroflexus Pigweed
Tidestromia lanuginosa Woolly Tidestromia

ANACARDIACEAE (Cashew Family)

Rhus trilobata

Skunkbush Sumac

Rhus microphylla Little-leaf Sumac

APOCYNACEAE (Dogbane Family)

Amsonia arenaria Blue-Star
Amsonia hirtella Blue-Star

ASCLEPIADACEAE (Milkweed Family)

Asclepias arenaria Sand Milkweed
Asclepias subverticillata

Poison Milkweed

ASTERACEAE (Composite Family)

TRIBE (2) AMBROSINEAE

Hymenoclea monogyra Singlewhirl Burro-bush

TRIBE (3) ASTEREAE

Baccharis bigelovii Bigelow Baccharis
Baccharis glutinosa Seepwillow Baccharis
Baccharis salicina Willow Baccharis
Chrysothamnus nauseosus

Rubber Rabbitbrush

Chrysothamnus pulchellus ssp. baileyi Southwest Rabbitbrush
Erigeron bellidastrum Fleabane
Gutierrezia microcephala Three Leaf Snakeweed
Gutierrezia sarothrae Broom Snakeweed
Haplopappus heterophyllus Jimmy-weed
Haplopappus spinulosus ssp. australis Goldenweed
Machaeranthera linearis Sand Goldenweed
Machaeranthera parviflora Wild Aster
Machaeranthera scabrella  
Machaeranthera tanacetifolia

Tohoka Daisy

TRIBE (5) HELIANTHEAE

Dicranocarpus parvilflorus Pitchfork
Engelmannia pinnatifida Engelmann Daisy
Flourensia cernua American Tarbush
Helianthus annuus Common Sunflower
Helianthus petiolaris Prairie Sunflower
Thelesperma megapotamicum Greenthread
Verbesina encelioides Golden Crownbeard
Zinnia grandiflora Rocky Mountain Zinnia

TRIBE (6) HELENIEAE

Bahia absinthifolia Bahia
Gaillardia pinnatifida Firewheel
Gaillardia pulchella Firewheel
Hymenopappus filifolius var. cinereus Yellow Cutleaf
Pectis angustifolia Lemonweed
Pseudoclappia arenaria False Clappia
Psilostrophe sparsiflora Paperdaisy
Psilostrophe tagetina Woolly Paperflower
Sartwellia flaveriae

Threadleaf Sartwell

TRIBE (8) SENECIONEAE

Senecio douglasii var.longilobus Threadleaf Groundsel
Senecio riddellii Riddell Groundsel
Senecio spartiodes Broom Groundsel

TRIBE (9) CYNAREAE

Centaurea melitensis Napa Star Thistle
Centaurea repens Russian Knapweed
Cirsium wheeleri Wheeler Thistle

TRIBE (10) MUTISIEAE

Perezia nana Desert Holly

BIGNONIACEA (Bignonia Family)

Chilopsis linearis

Desert Willow

BORAGINACEAE (Borage Family)

Coldenia hispidissima

Purple Borage

Cryptantha fulvocanescens Yellow Cryptantha
Heliotropium greggii Heliotrope
Lappula texana Stickseed

BRASSICACEAE (Mustard Family)

Descurainia pinnata Tansy-mustard
Dithyrea wislizenii Spectacle-Pod
Diplotaxus tenuifolia Slimleaf Wallrocket
Greggia camporum Velvety Greggia
Greggia camporam var. linearifolium

White Sands Mustard

Lepidium densiflorum Peppergrass
Lepidium montanum var. alyssoides

Pepperweed

Lesquerella fendleri Bladderpod
Sisymbrium irio London Rocket
Streptanthus validus Jewel Flower

CACTACEAE (Cactus Family)

Cereus greggii Night blooming Cereus
Coryphantha macromeris Nipple Beehive Cactus
Coryphantha sheeri Needle Beehive Cactus
Coryphantha vivipara var. vivipara Biscuit Cactus
Echinocactus horizonthalonius var. horizonthalonius Blue Barrel Cactus
Echinocereus fendleri var. fendleri Fendler's Hedgehog
Echinocereus fendleri var. rectispinus Fendler Needle-Spine Hedgehog
Echinocereus ennaecanthus var. stramineus Straw-colored Hedgehog
Echinocereus triglochidiatus var. gonacanthus

Claret Cup Hedgehog

Opuntia clavata Club Cholla
Opuntia imbricata Cane Cholla or Tree Cholla
Opuntia kleiniae var. kleiniae Klein Pencil Cholla
Opuntia leptocaulis

Desert Christmas Cactus

Opuntia macrorhiza var. macrorhiza Tuberous-Rooted Prickly Pear
Opuntia macrorhiza var. pottsii Plains Prickly Pear
Opuntia phaeacantha var. discata Desert Prickly Pear
Opuntia polyacantha var. rufispina Red-Spined Prickly Pear
Opuntia violacea var. macrocentra

Long-Spine Prickly Pear

Pediocactus papyracanthus Paper-Spine Plains Cactus

CHENOPODIACEAE (Goosefoot Family)

Allenrolfea occidentalis Pickle-weed
Atriplex canescens

Four-wing Saltbush

Salsola kali var. tenuifolia Russian Thistle
Suaeda depressa Seepweed
Suaeda suffrutescens Desert Seepweed
Suaeda torreyana Torrey Seepweed

CONVOLVULACEAE (Morning Glory Family)

Cressa truxillensis Silky Cressa

CUCURBITACEAE (Gourd Family)

Cucurbita foetidissima

Buffalo Gourd

EUPHORBIACEAE (Spurge Family)

Croton dioicus Croton
Euphorbia albomarginata Whitemargin Spurge
Euphorbia lata Hoary Euphorbia
Euphorbia serrula Sawtooth Spurge

FABACEAE (Pea Family)

Astragalus allorchrous Halfmoon Loco
Cassia bauhinioides Senna
Cassia lindheimeriana Senna
Dalea scoparia Broom Pea
Hoffmanseggia densiflora Hog-Potato
Melilotus officinalis Yellow Sweet Clover
Prosopis glandulosa

Honey Mesquite

FOUQUIERIACEAE (Ocotillo Family)

Fouquieria splendens

Ocotillo

FRANKENIACEAE (Frankenia Family)

Frankenia jamesii

Frankenia

GENTIANACEAE (Gentian Family)

Centaurium calycosum

Centaury or Rosita

Centaurium texense Lady Bird's Centaury
Eustoma exaltatum Catch-Fly Gentian
Eustoma grandiflorum Bluebell

HYDROPHYLLACEAE (Waterleaf Family)

Nama carnosum

Gyp Nama

Nama hispidum Hispid Nama
Phacelia corrugata Blue-Curls
Phacelia integrifolia

Scalloped Phacelia or Scorpionweed

KOEBERLINIACEAE (Junco Family)

Koeberlinia spinosa

Spiny All-thorn

LAMIACEAE (Mint Family)

Poliomintha incana

Hoary Rosemarymint

LOASACEAE (Stick-leaf Family)

Cevallia sinuata Cevellia
Mentzelia pumila var. integra

Blazingstar

Mentzelia pumila var. multiflora Desert Mentzelia
Mentzelia pumila var. pumila Stick-leaf

MALVACEAE (Mallow Family)

Sida leprosa var. depauperata Scrufy Sida
Sphaeralcea angustifolia Narrow-leaf Globemallow
Sphaeralcea incana

Soft Globemallow

Sphaeralcea subhastata Globemallow

NYCTAGINACEAE (Four O'clock Family)

Abronia angustifolia Sand Verbena
Allionia choisyi Smooth Trailing Four O'clock
Allionia incarnata Trailing Four O'clock
Ammocodon chenopodioides Goosefoot Moonpod
Mirabilis multiflora Colorado Four O'clock
Selinocarpus diffusus Spreading Moonpod
Selinocarpus lanceolatus Gyp Moonpod

ONAGRACEAE (Evening Primrose Family)

Calylophus hartwegii

Yellow Evening Primrose

Gaura coccinea Scarlet Gaura
Gaura parviflora Lizard Tail
Oenothera albicaulis Prairie Evening Primrose
Oenothera pallida runcinata White Evening Primrose
Oenothera pallida latifolia White Evening Primrose

PAPAVERACEAE (Poppy Family)

Argemone polyanthemos Prickly Poppy

PLUMBAGINACEAE (Leadwort Family)

Limonium limbatum

Sea-lavender

POLEMONIACEAE (Phlox Family)

Eriastrum diffusum Wooly Star
Ipomopsis pumila Low Gilia
Ipomopsis longiflora Pale Trumpets

POLYGONACEAE (Buckwheat Family)

Eriogonum rotundifolium Round Leaf Wild Buckwheat

RHAMNACEAE (Buck-thorn Family)

Condalia spathulata Knifeleaf Condalia

SALICACEAE (Willow Family)

Populus fremontii var. wizlizenii

Rio Grande Cottonwood

Salix gooddingii Goodding Willow

SANTALACEAE (Sandlewood Family)

Comandra pallida

Bastard-toadflax

SOLANACEAE (Potato Family)

Datura meteloides Jimson Weed
Datura querifolia Oak-Leaved Thornapple
Lycium berlandieri var. parviflorum Terrac Wolfberry
Lycium pallidum Pale Wolfberry
Lycium torreyi Torrey Wolfberry
Solanum elaeagnifolium

Silverleaf Nightshade

Solanum rostratum Buffalo Bur

TAMARICACEAE (Tamarix Family)

Tamarix gallica

Salt Cedar or Tamarisk

VERBENACEAE (Verbena Family)

Phyla incisa Texas Frog-fruit
Verbena bracteata Prostrate Vervain

VITACEAE (Grape Family)

Parthenocissus inserta Woodbine

ZYGOPHYLLACEAE (Caltrop Family)

Kallstroemia hirsutissima Carpetweed
Larrea tridentata Creosote Bush
Peganum harmala African Rue

SYNONOMY

For: See:
Andropus carnosus Nama carnosum
Aplopappus spinulosus Haplopapus spinulosus
Aster cichoriaceus Machaeranthera chichoriaceus
Aster linearis Machaeranthera linearis
Aster parvulus Machaeranthera parviflora
Aster tenacetifolius Machaeranthera tenacetifolia
Centaurea picris Centaurea repens
Chrysothamnus latisquameus Chrysothamnus nauseosus
Cladothrix lanigulosa Tidestroma lanuginosa
Comandra umbellata Commandra pallida
Crassina grandiflora Zinnia grandiflora
Cressa depressa Cressa truxillensis
Datura wrightii Datura meteloides
Dicranocarpus dicranocarpus Dicranocarpus parviflora
Distichlis spicata Distichlis stricta
Dimorphocarpa wislizenii Dithyrea wislizenii
Dondia moquini Suaeda torreyana
Dondia suffrutescens Suaeda suffrutescens
Erythrea texense Centaurium texense
Gilia pumila Ipomopsis pumila
Heterospermum dicranocarpum Dicranocarpus parviflora
Hoffmanseggia glauca Hoffmanseggia densiflora
Isocoma wrightii Haplopappus heterophyllus
Lepidium alyssoides Lepidium montanum
Lippia incisa Phyla incisa
Machaeranthera pinnatifida Haplopappus spinulosus
Nerisyrenia camporum Greggia camporum
Nerisyrenia linearifolia Greggia camporum var. linearifolia
Nuttallia procera Mentzelia pumila var. pumila
Oenothera hartwegii Calylophus hartwegii
Oenothera lavandulifolia Calylophus hartwegii
Oenothera runcinata Oenothera pallida runcinata
Opuntia engelmannii Opuntia phaeacantha discata
Phacelia corrugata Phacelia crenulata
Populus wizlizenii Populus fremontii var.wizlezenii
Rhus aromatica Rhus trilobata
Schizachyrium scoparium Andropogon scoparius
Scirpus maritimus Scirpus paludosus
Selinocarpus chenopodioides Ammocodon chenopodioides
Sida lepidota Sida leprosa
Sphaeralcea lobata Sphaeralcea angustifolia
Sporobolus asperifolia Muhlenbergia asperifolia
Sporobolus strictus Sporobolus contractus
Thelesperma gracile Thelesperma megapotamicum
Wootonia parviflora Dicranocarpus parviflorus
Xanthocephalum sarothrae Gutierrezia sarothrae
Yucca baccata var. torreyi Yucca torreyi

Last Updated: Wednesday, 22-Dec-2004 10:08:24 Eastern Standard Time
http://www.nps.gov/archive/whsa/plantlst.htm